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Sceloporus scalaris WIEGMANN, 1828

IUCN Red List - Sceloporus scalaris - Least Concern, LC

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Higher TaxaPhrynosomatidae, Sceloporinae; Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common NamesE: Light-bellied Bunch Grass Lizard
S: Lagartija Escamosa
E: Brown’s Bunchgrass Lizard [brownorum]
S: Escamosa de Brown [brownorum]
E: Southwestern Bunchgrass Lizard [unicanthalis]
S: Escamosa de Pastizal Sureña [unicanthalis] 
SynonymSceloporus scalaris WIEGMANN 1828: 370
Tropidolepis scalaris — DUMÉRIL & BIBRON 1837: 310
Tropidolepis Scalaris — DUMÉRIL, BIBRON & DUMÉRIL 1854: 271
Sceloporus scalaris — BOULENGER 1885: 234
Sceloporus scalaris scalaris — SMITH & TAYLOR 1950: 137
Sceloporus scalaris unicanthalis — SMITH & TAYLOR 1950: 138
Sceloporus scalaris — STEBBINS 1985: 125
Sceloporus scalaris — LINER 1994
Sceloporus scalaris scalaris — BELL et al. 2003 
DistributionMexico (C Durango, southward over the central and southern parts of the Mexican Plateau except (apparently) Guerrero. Distrito Federal, Guanajuato, Hidalgo, Jalisco, México, Michoacán, Puebla, Zacatecas.

Type locality: Mexico. Restricted to México, Distrito Federal, by SMITH & TAYLOR 1950. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous. 
TypesSyntypes: ZMB 654-656 (fide Taylor, 1969), collected by Ferdinand Deppe 
CommentTaxonomy: The S. scalaris species group includes five monotypic species, S. slevini, S. samcolemani, S. chaneyi, S. goldmani, and S. bicanthalis, and two polytypic species, S. scalaris (S. s. scalaris, S. s. unicanthalis, S. s. brownorum) and S. aeneus (S. a. aeneus, S. a. subniger).

Subspecies: Sceloporus scalaris brownorum, slevini, samcolemani and unicanthalis have been elevated to species status. 
EtymologyEtymology (scalaris): The Latin scalaris, "of a ladder," refers to the successive transverse dorsal dark lines resembling a ladder.

Etymology (brownorum): S. s. brownorum has been n amed for Dr. Bryce C. Brown, Director Emeritus of the Strecker Museum, Baylor University, Texas and his family, all of whom aided in collecting the largest series of this subspecies in any museum.

Etymology (unicanthalis): The Latin unus, "one," the Greek canthus, "corner," and the Latin suffix -alis, "pertaining to," refer to the distinctive single canthal scale. 
References
  • Ballinger, R. E.;Congdon, J. D. 1981. Population ecology and life history of a montane lizard (Sceloporus scalaris) in southeastern Arizona. Journal of Natural History 15: 213-222
  • Ballinger, R. E.;Congdon, J. D. 1996. Status of the bunch grass lizard, Sceloporus scalaris, in the Chiricahua Mountains of southeastern Arizona. Bull. Maryland Herp. Soc. 32: 67-69
  • Bell, E.L.; Smith, H.M. & Chiszar, D. 2003. AN ANNOTATED LIST OF THE SPECIES-GROUP NAMES APPLIED TO THE LIZARD GENUS SCELOPORUS. Acta Zoologica Mexicana (n.s.) 90: 103-174
  • Benabib, Miriam;Kjer, Karl M.;Sites, Jack W., Jr 1997. Mitochondrial DNA sequence-based phylogeny and the evolution of viviparity in the Sceloporus scalaris group (Reptilia, Squamata). Evolution 51 (4): 1262-1275 - get paper here
  • Blackburn, D.G. 1993. STANDARDIZED CRITERIA FOR THE RECOGNITION OF REPRODUCTIVE MODES IN SQUAMATE REPTILES. Herpetologica 49 (1): 118-132 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Bryson, Robert W.; Uri Omar García-Vázquez, Brett R. Riddle 2011. Relative roles of Neogene vicariance and Quaternary climate change on the historical diversification of bunchgrass lizards (Sceloporus scalaris group) in Mexico. Molecular Phylogenetics and Evolution 62 (1): 447-457 - get paper here
  • Carbajal-Márquez, Rubén A. and Gustavo E. Quintero-Díaz. 2013. Sceloporus scalaris (light-bellied bunchgrass lizard) clutch size. Herpetological Review 44 (4): 683
  • Carbajal-Márquez, Rubén A., Christian M. García-Balderas, Gustavo E. Quintero-Díaz, Ana G. Pérez-Delgadillo and Raramuri Reyes-Ardit. 2013. Sceloporus scalaris (light-bellied bunchgrass lizard) mating behavior. Herpetological Review 44 (4): 683-684
  • Castro Franco, R. & M. G. Bustos-Zagal 2003. Lagartijas de Morelos, México: distribución, hábitat y conservación. Acta Zoológica Mexicana (n.s.) 88:123-142
  • Castro-Franco, Rubén, María Guasalupe Bustos-Zagal 1994. List of reptiles of Morelos, Mexico, and their distribution in relation to vegetation types. Southwestern Naturalist 39 (2): 171-175 - get paper here
  • Cobarrubias, Edgar Flores; Daniel Cruz-Sáenz, and David Lazcano 2012. Notes on the Herpetofauna of Western Mexico 6: Amphibians and Reptiles of Hostotipaquillo, Jalisco, Mexico. Bull. Chicago Herp. Soc. 47 (2):21-26 - get paper here
  • Creer, Douglas A.;Kjer, Karl M.;Simmons, Daniel L.;Sites, Jack W., Jr 1997. Phylogenetic relationships of Sceloporus scalaris species group (Squamata). Journal of Herpetology 31 (3): 353-364 - get paper here
  • Degenhardt, William G.; C. W. Painter, and A. H. Price 1996. Amphibians and reptiles of New Mexico. Univ. New Mexico Press, 431 pp.
  • Dixon, James R. and Julio A. Lemos-Espinal 2010. Amphibians and reptiles of the state of Queretaro, Mexico. Tlalnepantla UNAM, 428 pp.
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Duméril, A. M.C., G. BIBRON & A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S. - get paper here
  • Fitch, H. S. 1978. Sexual size differences in the genus Sceloporus. Univ. Kansas Sci. Bull. 51 (13): 441-461 - get paper here
  • Gravenhorst, J. L. C. 1838. Beiträge zur genaueren Kenntniss einiger Eidechsengattungen. Nova Acta Acad. Caes. Leop.-Carol. 18: 712-784 [1837]
  • GRUMMER, JARED A. & ROBERT W. BRYSON JR. 2014. A new species of bunchgrass lizard (Squamata: Phrynosomatidae) from the southern sky islands of the Sierra Madre Occidental, Mexico. Zootaxa 3790 (3): 439–450 - get paper here
  • Howland, Jeffrey M.;Whittinghill-Howland, Jan M. 1995. Sceloporus scalaris. Herpetological Review 26 (3): 155 - get paper here
  • Leaché, A.D. 2010. Species trees for spiny lizards (Genus Sceloporus): Identifying points of concordance and conflict between nuclear and mitochondrial data. Molecular Phylogenetics and Evolution 54: 162–171 - get paper here
  • Leaché, A.D. & J.W.Sites, Jr. 2009. Chromosome Evolution and Diversification in North American Spiny Lizards (Genus Sceloporus). Cytogenet Genome Res 127: 166-181 - get paper here
  • Lemos-Espinal, Julio A. and James R. Dixon 2013. Amphibians and Reptiles of San Luis Potosí. Eagle Mountain Publishing, xii + 300 pp.
  • Lemos-Espinal, Julio A., Geoffrey R. Smith 2015. Amphibians and reptiles of the state of Hidalgo, Mexico. Check List 11 (3): 1642 - get paper here
  • Leyte-Manrique, Adrian; Efrén M. Hernández Navarro y Luis A. Escobedo-Morales 2015. Herpetofauna de Guanajuato: Un análisis histórico y contemporáneo de su conocimiento Revista Mexicana de Herpetología 1(1): 1–14 - get paper here
  • Martin, Plul S. 1958. A biogeography of reptiles and amphibians in the Gomez Farias Region, Tamaulipas, Mexico. Miscellaneous publications, Museum of Zoology, University of Michigan (101): 1-102 + 7 plates - get paper here
  • Mink, Daniel G.;Sites, Jack W., Jr 1996. Species limits, phylogenetic relationships, and origins of viviparity in the scalaris complex of the lizard genus Sceloporus (Phrynosomatidae; Sauria). Herpetologica 52 (4): 551-571 - get paper here
  • Muñoz-Nolasco, Francisco Javier; Daniel Cruz-Sáenz, Oscar Jair Rodríguez-Ruvalcaba and Isaac Enrique Terrones-Ferreiro 2015. Notes on the Herpetofauna of Western Mexico 12: Herpetofauna of a Temperate Forest in Mazamitla, Southeastern Jalisco, Mexico. Bull. Chicago Herp. Soc. 50 (4):45-50
  • Ortega, Alfredo; Ricardo Rodriguez, Lucina Hernandez and Robert Barbault. 1984. Cycles Journaliers d’Activité chez deux Espèces Sympatriques de Sceloporus (Iguanidae), S. grammicus et S. scalaris Amphibia-Reptilia 5 (3-4): 347 - 354 - get paper here
  • Smith H M.; Watkins-Colwell G J.; Lemos-Espinal J A.; Chiszar D. 1997. A new subspecies of the lizard (Sceloporus scalaris (Reptilia: Sauria: Phrynosomatidae) from the Sierra Madre Occidental of Mexico. Southwestern Naturalist 42 (3): 290-301. - get paper here
  • Smith H.M.; Bock C.E.; Bock J.H., 1990. Notes on reproduction and coloration in the bunchgrass lizard Sceloporus scalaris in southeastern Arizona USA. Bull. Maryland Herp. Soc.: 64-67
  • Smith, H. M. 1939. The Mexican and Central American lizards of the genus Sceloporus. Field Mus. Nat. Hist., Zool. Ser. 26: 1-397. - get paper here
  • Smith, H. M.;Hall, W. P. 1974. Contributions to the concepts of reproductive cycles and the systematics of the scalaris group of the lizard genus Sceloporus. Great Basin Naturalist 34 (2): 97-104 - get paper here
  • Smith,H.M. 1937. A synopsis of the Scalaris group of the lizard genus Sceloporus. Occ. Pap. Mus. Zool. Univ. Mich 361: 1-8
  • Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston
  • Terán-Juárez, Sergio A., Elí García Padilla, Vicente Mata-Silva, Jerry D. Johnson and Larry David Wilson. 2016. The herpetofauna of Tamaulipas, Mexico: composition, distribution, and conservation status. Mesoamerican Herpetology 3 (1): 43–113 - get paper here
  • Thomas, Robert A. & Dixon,J.R. 1976. A re-evaluation of the Sceloporus scalaris group (Sauria: Iguanidae). Southwestern Naturalist 20 (4): 523-536 - get paper here
  • Valdez-Lares, R.; R. Muñiz-Martínez; E.Gadsden; G. Aguirre-León; G. Castañeda-Gaytán; R. Gonzalez-Trápaga 2013. Checklist of amphibians and reptiles of the state of Durango, México. Check List 9 (4):714-724 - get paper here
  • Vázquez Díaz, Joel;Quintero Díaz, Gustavo E. 2005. Anfibios y Reptiles de Aguascalientes [2nd ed.]. CONABIO, CIEMA, 318 pp.
  • Webb, R.G. 1984. Herpetogeography in the Mazatlán-Durango Region of the Sierra Madre Occidental, Mexico. Vetrebrate Ecology and Systematics - A ribute to Henry S. Fitch; Museum of Natural History, University of Kansas, Lawrence, pp. 217-241
  • Webb, Robert G.; Smith, Hobart M.; Chiszar, David; Watkins-Colwell, Gregory J. 1996. Geographic Distribution. Sceloporus scalaris unicanthalis. Herpetological Review 27 (3): 153 - get paper here
  • Wiegmann,A.F.A. 1828. Beyträge zur Amphibienkunde. Isis von Oken 21 (4): 364-383 - get paper here
 
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