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Sceloporus squamosus BOCOURT, 1874

IUCN Red List - Sceloporus squamosus - Least Concern, LC
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Higher TaxaPhrynosomatidae, Sceloporinae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Mexican Spiny Lizard, Dwarf Spiny Lizard
S: Espinosa Enana 
SynonymSceloporus squamosus BOCOURT 1874: 212
Sceloporus fulvus BOCOURT 1874: 214 (fide SMITH 1939)
Sceloporus squamosus — BOULENGER 1885: 238
Sceloporus fulvus — BOULENGER 1885: 238
Sceloporus squamosus — SMITH 1939: 319
Sceloporus squamosus — SMITH & TAYLOR 1950: 135
Sceloporus squamosus — LINER 1994
Sceloporus squamosus — KÖHLER 2000: 85
Sceloporus squamosus — WIENS et a. 2010
Sceloporus squamosus — MCCRANIE 2018
Sceloporus squamosus — HEIMES 2022 
DistributionMexico (Oaxaca, E Chiapas), Guatemala, El Salvador, Nicaragua, Costa Rica, Honduras

fulvus: Type Locality: "Environs de Guatemala et de l'Antigua". La Unión, El Salvador, fide Guibe
(1954).

Type locality: “Environs de Guatemala et de l’Antigua” [= Volcán Antigua], 1500 m elevation.  
Reproductionoviparous 
TypesSyntypes: MNHN-RA 3180, MNHN-RA 3180A-C, MNHN-RA 3181, MNHN-RA 3181A, MNHN-RA 3182, MNHN-RA 3182A (8 specimens), USNM 10964, collected by Bouvier
Syntypes: MNHN 3179(6) [fulvus] 
DiagnosisDiagnosis: Smith 1939: 319 
CommentNot listed for Mexico by KÖHLER (2000). 
EtymologyThe Latin noun squamosus, "scaly," refers to the quite large scales for such a small lizard.

fulvus: This Latin noun fulvus, means "tawny, reddish yellow," and refers to the dark reddish-brown color of both sexes. 
References
  • Bocourt, M. E. 1874. Etudes sur les Reptiles. Mission Scientifique au Mexique et dans l'Amérique Centrale. Recherches Zoologiques.Vol. 3 (sect. 1). Imprimerie Impériale, Paris. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Brongersma, L. D. 1954. On some lizards from the Republic of El Salvador. Konikl. Nederl. Akad. van Wetenschappen (Amsterdam) 57 (2): 165-174
  • Casas-Andreu, G., F.R. Méndez-De la Cruz and X. Aguilar-Miguel. 2004. Anfibios y Reptiles; pp. 375–390, in A.J.M. García-Mendoza, J. Ordoñez and M. Briones-Salas (ed.). Biodiversidad de Oaxaca. Instituto de Biología, UNAM-Fondo Oaxaqueño para la Conservación de la Naturaleza-World Wildlife Fund, México, D. F.
  • Eisenberg, T. & G. Köhler 1996. Lebensweise, Pflege und Nachzucht von Sceloporus squamosus Bocourt, 1874. Sauria 18 (1): 11-14 - get paper here
  • Gutsche, Alexander 2015. Die Insel der verschollenen Salamander: Zur Herpetofauna der honduranischen Isla del Tigre. Terraria-Elaphe 2015 (3): 38-43 - get paper here
  • Heimes, P. 2022. LIZARDS OF MEXICO - Part 1 Iguanian lizards. Edition Chimaira, Frankfurt Am Main, 448 pp.
  • Johnson, Jerry D.; Vicente Mata-Silva, Elí García Padilla, and Larry David Wilson 2015. The Herpetofauna of Chiapas, Mexico: composition, distribution, and conservation. Mesoamerican Herpetology 2 (3): 272–329. - get paper here
  • Köhler, G. 2000. Reptilien und Amphibien Mittelamerikas, Bd 1: Krokodile, Schildkröten, Echsen. Herpeton Verlag, Offenbach, 158 pp.
  • Köhler, G. 2008. Reptiles of Central America. 2nd Ed. Herpeton-Verlag, 400 pp.
  • Köhler, G. & Heimes, P. 2002. Stachelleguane. Herpeton-Verlag (Offenbach), 174 pp.
  • Köhler, Gunther, Fabian Schmidt and Pascal Eusemann 1998. Morphologie, Verbreitung und Fortpflanzung der Stachelschuppenleguane in El Salvador: 1. Sceloporus squamosus Bocourt, 1874. Salamandra 34 (4): 301-308 - get paper here
  • Leenders, T. 2004. Der Nationalpark “El Imposible” in El Salvador. Reptilia (Münster) 9 (48): 45-49 - get paper here
  • Martínez-Fonseca JG, Holmes IA, Sunyer J, Westeen EP, Grundler MR, Cerda PA, Fernández-Mena MA, Loza-Molina JC, Monagan IV Jr., Nondorf D, Pandelis GG, Rabosky ARD 2023. A collection and analysis of amphibians and reptiles from Nicaragua with new country and departmental records. Check List 20(1): 58-125 - get paper here
  • Mata-Silva, Vicente, Jerry D. Johnson, Larry David Wilson and Elí García-Padilla. 2015. The herpetofauna of Oaxaca, Mexico: composition, physiographic distribution, and conservation status. Mesoamerican Herpetology 2 (1): 6–62 - get paper here
  • McCranie, James R. 2015. A checklist of the amphibians and reptiles of Honduras, with additions, comments on taxonomy, some recent taxonomic decisions, and areas of further studies needed. Zootaxa 3931 (3): 352–386 - get paper here
  • McCranie, James R. 2018. The Lizards, Crocodiles, and Turtles of Honduras. Systematics, Distribution, and Conservation. Bulletin of the Museum of Comparative Zoology, Special Publication Series (2): 1- 666 - get paper here
  • McCranie, James R., and Alexander Gutsche 2016. The herpetofauna of islands in the Golfo de Fonseca and adjacent waters, Honduras. Mesoamerican Herpetology 3 (4): 842–899 - get paper here
  • Mertens, R. 1952. Die Amphibien und Reptilien von El Salvador. Abh. senckenb. naturf. Ges. (Frankfurt) (No. 487): 120 pp.
  • Savage, J.M. 2002. The Amphibians and Reptiles of Costa Rica: A Herpetofauna Between Two Continents, Between Two Seas. University of Chicago Press, 934 pp. [review in Copeia 2003 (1): 205]
  • Smith, H. M. 1939. The Mexican and Central American lizards of the genus Sceloporus. Field Mus. Nat. Hist., Zool. Ser. 26: 1-397. - get paper here
  • Smith, H.M. & Taylor,E.H. 1950. An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bull. US Natl. Mus. 199: 1-253 - get paper here
  • Solís, J. M., L. D. Wilson, and J. H. Townsend. 2014. An updated list of the amphibians and reptiles of Honduras, with comments on their nomenclature. Mesoamerican Herpetology 1: 123–144 - get paper here
  • Stuart, L.C. 1963. A checklist of the herpetofauna of Guatemala. Misc. Publ. Mus. Zool., Univ. Michigan (No. 122): 1-150 - get paper here
  • Sunyer, Javier 2014. An updated checklist of the amphibians and reptiles of Nicaragua. Mesoamerican Herpetology 1 (2): 186–202. - get paper here
  • Sunyer, Javier & José́ Gabriel Martínez-Fonseca 2023. An updated country checklist to the amphibians and reptiles of Nicaragua. REVISTA NICARAGÜENSE DE BIODIVERSIDAD (100): 1-25 - get paper here
  • Vences M., Franzen M., Flaschendräger A., Schmitt R. & Regös J. 1998. Beobachtungen zur Herpetofauna von Nicaragua: kommentierte Artenliste der Reptilien. Salamandra 34 (1): 17-42 - get paper here
  • Wiens, John J.; Caitlin A. Kuczynski, Saad Arif & Tod W. Reeder 2010. PHYLOGENETIC RELATIONSHIPS OF PHRYNOSOMATID LIZARDS BASED ON NUCLEAR AND MITOCHONDRIAL DATA, AND A REVISED PHYLOGENY FOR SCELOPORUS. Molecular Phylogenetics and Evolution 54: 150-161 - get paper here
 
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