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Sceloporus torquatus WIEGMANN, 1828

IUCN Red List - Sceloporus torquatus - Least Concern, LC

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Higher TaxaPhrynosomatidae, Sceloporinae, Iguania, Sauria, Squamata (lizards)
Common NamesE: Crevice Swift, Torquate Lizard, Wiegmann's Torquate Lizard
S: Espinosa de Collar 
SynonymSceloporus torquatus WIEGMANN 1828: 369
Agama torquata PEALE & GREEN 1830
Tropidolepis torquatus — DUMÉRIL & BIBRON 1837: 301
Tropidolepis Torquatus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 270
Sceloporus ferrariperezi COPE 1885: 400
Sceloporus torquatus torquatus — COPE 1885: 402
Sceloporus torquatus — BOULENGER 1885: 219
Sceloporus ferrariperezi ferrariperezi — SMITH 1939: 214
Sceloporus ferrariperezi ferrariperezi — SCHMIDT 1947
Sceloporus torquatus torquatus — SMITH & TAYLOR 1950: 125
Sceloporus torquatus — LINER 1994
Sceloporus torquatus — WIENS et a. 2010
Sceloporus torquatus — CAMPILLO-GARCÍA et al. 2021
Sceloporus torquatus — CORTÉS-VÁZQUEZ et al. 2022 
DistributionMexico (Nayarit and other states [HR 30: 111], Durango, Guerrero, Tamaulipas)

torquatus: C Mexico, including Hidalgo, SC Veracruz, México, Distrito Federal, N Puebla, E Morelos, S Guanajuato, N Michoacán. Type locality: Mexico. Restricted to México, Distrito Federal, by SMITH & TAYLOR 1950.

ferrariperezi: Mexico (Guanajuato, Michoacan); Type Locality: Guanajuato, Guanajuato, Mexico.  
Reproductionviviparous (all species of the torquatus group are viviparous) 
TypesLectotype: ZMB 629, paralectotype: ZMB 631; some of the former syntypes (ZMB 628-631), collected by Ferdinand Deppe, belong to other species, see comments.
Holotype: ANSP 8499; Mexico [Agama torquata]
Syntypes: USNM 9874, 9876, 9878, 9880, 9895 (latter exchanged to MCZ in 1944 fide Cochran, 1961) [ferrariperezi]. 
DiagnosisDefinition (genus): genus with femoral pores, depressed body, imbricate and keeled dorsal scales, enlarged occipital shield, keeled digital lamellae, and distinct tympanum, in combination with the absence of abdominal ribs, collar, gular pouch, pterygoid teeth, and dorsal crest, distinguished from the other genera of the family (Smith 1939: 28). 
CommentType species: Sceloporus torquatus is the type species of the genus Sceloporus Wiegmann, 1828. For a recent phylogeny of the genus see WIENS et al. 2010.

Subspecies: Sceloporus torquatus minor and other subspecies are now treated as full species or as a subpsecies of S. jarrovii.

Synonymy: Agama torquata PEALE & GREEN 1830 has been described independently and given (serendipitoulsy!) the same species epithet. The name is thus a junior secondary homonym and objective synonym of Sceloporus torquatus Wiegmann (1828).

Sympatry: Circular sympatry may occur between S. t. mikeprestoni, derived from S. t. torquatus, and S. t. binocularis, derived from S. t. melanogaster. Additional field work in the hiatus between the ranges of S. t. mikeprestoni and S. t. binocularis is necessary to resolve this problem (BELL et al. 2003) but see CAMPILLO-GARCÍA et al. 2021 for an update.

Types: One of the syntypes of Sceloporus ferrariperezi COPE (USNM 9880) is actually Sceloporus spinosus according to SMITH & TAYLOR 1950. One of the syntypes of S. torquatus, ZMB 628 was determined to be Sceloporus aureolus Smith, 1942 by Campillo-Garcia et al. 2021. These authors also re-determined the specimen ZMB 630, another syntype of S. torquatus, as S. t. melanogaster by having undivided supraocular scales, 30 dorsal scales, 41 ventral scales, diffuse dark nuchal collar interrupted by dorsolateral light bands or marks, as well as a series of dark irregular spots that fade over the base of the tail. Finally, they also found that specimen ENCB 5756, a paratype of S. t. mikeprestoni, actually pertains to Sceloporus minor Cope, 1885. This specimen has divided supraocular scales, 36 dorsal scales, 40 scales around the body, and 44 ventral scales.

Group: S. torquatus group (sensu lato, fide Flores-Villela et al. 2022). The torquatus group consists of 8 species fide Flores-Villela et al. 2022: S. binocularis, S. bul- leri Boulenger, S. insignis Webb, S. jarrovii Cope, S. madrensis, S. melanogaster, S. mikeprestoni and S. torquatus.

Distribution: Not in Querétaro fide Tepos-Ramírez et al. (2023). 
EtymologyNamed after the Latin torquatus, "adorned with a necklace or collar", refers to the conspicuous, light-bordered black collar in these lizards. 
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