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Scincella modesta (GÜNTHER, 1864)

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
SubspeciesScincella modesta modesta (GÜNTHER 1864)
Scincella modesta septentrionalis (SCHMIDT 1925) 
Common NamesE: Modest Ground Skink
Chinese: 宁波滑蜥 
SynonymEumeces modestus GÜNTHER 1864: 87
? Mocoa exigua ANDERSON 1878
Lygosoma laterale — BOULENGER 1887
Leiolopisma laterale reevesii — VAN DENBURGH 1912
Leiolopisma laterale formosensis VAN DENBURGH 1912
Leiolopisma monticola SCHMIDT 1927: 497 (fide OUBOTER 1986)
Leiolopisma septentrionale SCHMIDT 1927
Leiolopisma modestum SCHMIDT 1927
Leiolopisma reevesii — POPE 1935
Leiolopisma modestum — SMITH 1935
Scincella reevesii — WEBB, JONES & BYERS 1956
Scincella modesta MITTLEMAN 1952
Leiolopisma tsinlingensis HU & DJAO 1966
Scincella monticola — GREER 1974
Scincella modesta — GREER 1974: 7
Scincella septentrionalis — GREER 1974
Scincella tsinlingensis — GREER 1974
Scincella formosensis — GREER 1974
Scincella exigua — GREER 1974
Scincella modesta — OUBOTER 1986: 51
Scincella modesta modesta — WANG & ZHAO 1986
Scincella modesta — LINKEM, DIESMOS & BROWN 2011
Scincella modesta — WANG et al. 2022

Scincella modesta septentrionalis (SCHMIDT 1925)
Leiolopisma septentrionalis SCHMIDT 1925: 2
Scincella septentrionalis — SHEA & GREER 2002: 156 
DistributionC China (northward to Hebei in the east, Fujian, Zhejiang)

Type locality: Ningpo (= Ningbo), Zhejiang Province, China.  
Reproductionoviparous 
TypesSyntypes: BMNH 1946.8.16.75-76
Holotype: CAS 21678 [boettgeri]
Holotype: MCZ 7966 [schmidti]
Holotype: ZSI 4270 [Mocoa exigua Anderson 1879]
Holotype: AMNH 21451 [septentrionalis] 
DiagnosisDiagnosis: A moderately sized Scincella. Body slightly elongated. No projecting lobules. Small number of subdigital lamellae under the fourth toe. Lateral dark band distinct, usually not bordered by a lighter dorsolateral stripe and not broken up by lighter spots (Ouboter 1986: 52). 
CommentSee discussion about synonyms in Ouboter (1986) who doesn’t distinguish between subspecies. Scincella vandenburghi (SCHMIDT 1927) and S. boettgeri have been removed from synonymy of S. modesta. S. potanini and S. schmidti have been considered as synonyms of S. modesta but are listed as valid species here.

The original description is available online (see link below). 
References
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • Chen, Szu-Lung; Tsutomu Hikida; Han, Sang-Hoon; Shim, Jae-Han; Oh, Hong-Shik & Hidetoshi Ota 2001. Taxonomic status of the Korean populations of the genus Scincella (Squamata: Scincidae). Journal of Herpetology 35 (1): 122-129 - get paper here
  • Greer, A.E. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology 31: 1-67. - get paper here
  • Günther, A. 1864. The Reptiles of British India. London (Taylor & Francis), xxvii + 452 pp. - get paper here
  • Lazell, James; Lau, Michael;Lu, Wenhua 1997. A brief herpetological excursion to Wai Ling Ding, Wanshan Islands, South China Sea. Asiatic Herpetological Research 7: 80-84 - get paper here
  • Linkem, Charles W.; Arvin C. Diesmos, Rafe M. Brown 2011. Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163: 1217–1243 - get paper here
  • Ouboter P E 1986. A revision of the genus Scincella (Reptilia: Sauria: Scincidae) of Asia, with some notes on its evolution. Zoologische Verhandelingen (229): 1-66 - get paper here
  • Pope, CLIFFORD H. 1935. The Reptiles of China. Turtes, Crocodilians, Snakes, Lizards. Amer. Mus. Nat. Hist., New York, Nat. Hist. Central Asia, 10: lii, 1-604 - get paper here
  • Schmidt, K.P. 1927. Notes on Chinese reptiles. Bull. Amer. Mus. Nat. Hist., New York, 54 (4): 467-551 - get paper here
  • Schmidt, KARL P. 1925. New reptiles and a new salamander from China. American Museum Novitates (157): 1-5. - get paper here
  • Shea, Glenn M. and Allen E. Greer 2002. From Sphenomorphus to Lipinia: Generic reassignment of two poorly known New Guinea skinks. Journal of Herpetology 36 (2): 148-156 - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Van Denburgh, J., 1912. Concerning certain species of reptiles and amphibians from China, Japan, the Loo Choo Islands, and Formosa. Proc. Cal. Ac. Sci. (Series 4) 3 (10): 187-258. - get paper here
  • Wang, C. G. M. Z. X., Jia, S. Y. K. T. L., & Hu, C. Z. J. 2022. An updated species checklist of amphibians and reptiles in Fujian Province, China. Biodiversity Science, 22090 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Wang, YUE-ZHAO 1986. Studies on Chinese species of Scincella (Scincidae, Sauria) [in Chinese]. Acta Herpetol. Sinica, Chengdu, [new ser.], 5 (4): 267-277 - get paper here
  • Yang, Jing; Yan-Yan Sun, Tian-Boa Fu, Da-De Xu, Xiang Ji 2012. Selection for increased maternal body volume does not differ between two Scincella lizards with different reproductive modes. Zoology 115 (4): 199-206 - get paper here
  • Zhang, Minchu; Chaonan Tang, Qing Zhang, Chengxiu Zhan, Chuanwu Chen, Yanping Wang 2024. Selective extinction and habitat nestedness are the main drivers of lizard nestedness in the Zhoushan Archipelago. Current Zoology, 70 (2): 244–252, - get paper here
 
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