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Sphenomorphus tonkinensis NGUYEN, SCHMITZ, NGUYEN, ORLOV, BÖHME & ZIEGLER, 2011

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Higher TaxaScincidae, Sphenomorphinae, Scincoidea, Sauria, Squamata (lizards) 
Subspecies 
Common NamesE: Tonkin Forest Skink
G: Tonkin-Bodenskink 
SynonymSphenomorphus tonkinensis NGUYEN, SCHMITZ, NGUYEN, ORLOV, BÖHME & ZIEGLER 2011
Sphenomorphus rufocaudatus — SCHMITZ & ZIEGLER 2003 (not DAREVSKY & NGUYEN 1983) 
DistributionVietnam

Type locality: within Cat Ba National Park, Cat Ba Island, Hai Phong, (20°47.610’N, 106°59.462’E), elevation 100 m. Map legend:
Type locality - Type locality.
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproduction 
TypesHolotype: IEBR A.0730, adult male, collected on 19 May 2007 by T. Q. Nguyen, R. Stenke, and T. Ziegler.
Paratypes.—ZFMK 61734, adult male, collected in April 1995 by P. Heimes from Wuzhishan, Hainan Island, China, 700 m a.s.l. (previously reported as Sphenomorphus rufocaudatus by Schmitz and Ziegler, 2003); ZFMK 82864, subadult, collected by G. Vogel from Tam Dao Mt., Vinh Phuc Province (unknown collection date); five specimens from Ba Mun Island (21u04.3679N, 107u35.6339E) and one specimen (VNMN 949) from Cai Lim Island (21u06.5009N, 107u34.7009E) within Bai Tu Long National Park, Quang Ninh Province, in the Gulf of Tonkin: VNMN 205, adult male, collected on 20 September 2008 by T. T. Nguyen and S. Ryabov; IEBR A.0912 and MHNG 2713.33, adult males, collected on 16 May 2009 by T. Ziegler and T. Q. Nguyen; VNMN 934, 935, 949, adult females, collected by T. T. Nguyen from 21–23 September 2009. 
CommentS. tonkinensis is differentiated from the congeners by the combination of the following characters: size small (SVL 35.8–48.8 mm); prefrontals in contact with each other mesially; supralabials seven; primary temporals two; external ear opening present, without lobules, tympanum slightly sunk; midbody scales in 32–34 rows; dorsal scales smooth, paravertebral scales 65–72, not widened; limbs well developed, pentadactyl; 15–19 smooth lamellae under fourth toe; free margins of upper and lower eyelids edged in white; neck, dorsum, and tail base bronze-brown with a discontinuous dark vertebral stripe; upper lateral zone black, interrupted by small light spots from behind the neck.

Diagnosis.—Size small (SVL 35.8–48.8 mm); prefrontals in contact with each other mesially; supraoculars four; a pair of nuchals present or absent; lower eyelid moveable, scaly; supralabials seven; primary temporals two; external ear present, without lobules, tympanum slightly sunk; midbody scales in 32–34 rows; dorsal scales smooth, paravertebral scales 65–72, not widened; limbs well developed, pentadactyl; scales on dorsal surface of base of digit IV in three rows; 15–19 smooth lamellae under fourth toe; free margins of upper and lower eyelids edged in white; dorsum and tail base bronze brown with a discontinuous dark vertebral stripe; upper lateral zone with black stripe, interrupted by small light spots from behind the neck. 
EtymologyEtymology.—The specific epithet tonkinensis refers to the former name of North Vietnam, Tonkin, and also refers to the Gulf of Tonkin (Vinh Bac Bo), which houses the type locality. 
References
  • Gawor, A., C. T. Pham, T. Q. Nguyen, T. T. Nguyen, A. Schmitz & T. Ziegler 2016. The herpetofauna of the Bai Tu Long National Park, northeastern Vietnam. Salamandra 52 (1): 23-41 - get paper here
  • Hecht, Vera L.; Cuong T. Pham, Tao T. Nguyen, Truong Q. Nguyen, Michael Bonkowski & Thomas Ziegler 2013. First report on the herpetofauna of Tay Yen Tu Nature Reserve, northeastern Vietnam. Biodiversity Journal 4 (4): 507–552
  • Nguyen, Truong Quang; Andreas Schmitz, Tao Thien Nguyen, Nikolai L. Orlov, Wolfgang Böhme, and Thomas Ziegler 2011. Review of the Genus Sphenomorphus Fitzinger, 1843 (Squamata: Sauria: Scincidae) in Vietnam, with Description of a New Species from Northern Vietnam and Southern China and the First Record of Sphenomorphus mimicus Taylor, 1962 from Vietnam. Journal of Herpetology 45 (2): 145-154. - get paper here
  • ZIEGLER, T., RAUHAUS, A., TRAN, T.D., PHAM, C.T., VAN SCHINGEN, M., DANG, P.H., LE, M.D. & NGUYEN, T.Q. 2015. Die Amphibien- und Reptilienfauna der Me-Linh-Biodiversitätsstation in Nordvietnam Sauria 37 (4): 11-44 - get paper here
 
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