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Stegonotus derooijae RUANE, RICHARDS, MCVAY, TJATURADI, KREY & AUSTIN, 2017

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common Names 
SynonymStegonotus derooijae RUANE, RICHARDS, MCVAY, TJATURADI, KREY & AUSTIN 2017: 13
Stegonotus derooijae — KAISER et al. 2018: 56 
DistributionIndonesia (West Papua: Raja Ampat Islands of Batanta, Salawati and Waigeo)

Type locality: Waibya Camp, Salawati Island, Raja Ampat Regency, West Papua, Indonesia, elevation 75 m (0.956383°S, 130.784333°E).  
Reproduction 
TypesHolotype: MZB Ophi.3288 (field number SJR 7788, tissue ABTC 90225), adult male. Collected by S. Richards, B. Tjaturadi and K. Krey on 26 June 2005. Paratypes: MZB.Ophi.3293 (field number SJR 7789, tissue ABTC 90226), same data as holotype; SAMA R70467 (field number SJR 7586, tissue ABTC 90123) collected by S. Richards, B. Tjaturadi and K. Krey on 9 June 2005 from Warinkabom, Batanta Island, Raja Ampat Regency, West Papua Province, Indonesia, elevation 50 m asl (0.836942°S, 130.72162°E). 
DiagnosisA new species of Stegonotus that can be diagnosed from its congeners by the following combination of characters: a dark grey or brown unpatterned dorsum with a white or cream venter with dorsal scale pigmentation continuing onto some or many of the ventral scales, dark grey pigmentation on the subcaudal scales extending from the dorsal scale pigmentation and with grey on the anterior edges of some subcaudal scales, 17 dorsal scale rows at midbody, 178−197 ventral scales and 90−94 divided subcaudal scales, two preocular scales, seven (rarely nine) supralabial scales, eight or nine infralabial scales, with the supralabial and infralabial scales cream and with grey pigmentation.

Comparisons: Stegonotus derooijae sp. nov. is part of the S. diehli complex (Figure 3) and this taxon would have been formerly considered S. diehli. It differs morphologically from S. diehli by having 17 dorsal scale rows at midbody versus 15 dorsal scale rows in S. diehli. Stegonotus derooijae sp. nov. also differs from S. diehli in having higher numbers of ventral scales (178−197 in S. derooijae sp. nov. versus 162−174 in S. diehli) and subcaudal scales (90−94 in S. derooijae versus 73−85 in S. diehli) for the specimens we examined (excluding specimens with incomplete tails; Appendix 1). It differs from the new species (described below) from Southern Highlands and Chimbu Provinces by having two rather than (usually) one preocular scale. The overall dark coloration of S. derooijae sp. nov. distinguishes it from the sympatric S. iridis sp. nov. described above.

Comparison: For a comprehensive comparison of species, especially to the new species Stegonotus ayamaru, see that entry and Kaiser et al. 2019. 
CommentHabitat: forest floor in moderately to heavily disturbed lowland (< 100 m above sea level) rainforest.

Sympatry: S. iridis 
EtymologyThe species epithet is a noun in the genitive case and is a patronym in honour of Dr Nelly De Rooij, who described many taxa from New Guinea and Indonesia, including Stegonotus florensis. 
References
  • Kaiser, Christine M.; Hinrich Kaiser & Mark O'Shea 2018. The taxonomic history of Indo-Papuan groundsnakes, genus Stegonotus Duméril et al., 1854 (Colubridae), with some taxonomic revisions and the designation of a neotype for S. parvus (Meyer, 1874) Zootaxa 4512 (1): 001–073
  • KAISER, CHRISTINE M.; MARK O’SHEA, HINRICH KAISER 2019. A new species of Indo-Papuan groundsnake, genus Stegonotus Duméril et al., 1854 (Serpentes, Colubridae), from the Bird’s Head Peninsula of West Papua, Indonesia, with comments on differentiating morphological characters Zootaxa 4590 (2): 201-230 - get paper here
  • Ruane, S., Richards, S. J., McVay, J. D., Tjaturadi, B., Krey, K., & Austin, C. C. 2017. Cryptic and non-cryptic diversity in New Guinea ground snakes of the genus Stegonotus Duméril, Bibron and Duméril, 1854: a description of four new species (Squamata: Colubridae). Journal of Natural History 52: 917-944 - get paper here
 
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