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Stenocercus nigrobarbatus VENEGAS, ECHEVARRÍA, GARCÍA-AYACHI & LANDAURO, 2020

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Higher TaxaTropiduridae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common Names 
SynonymStenocercus nigrobarbatus VENEGAS, ECHEVARRÍA, GARCÍA-AYACHI & LANDAURO 2020 
DistributionPeru (Huancavelica)

Type locality: Fundición (12°17’36.41”S, 74°39’39.67” W) at 2,158 m asl, Tayacaja Province, Huancavelica Region, Peru  
Reproduction 
TypesHolotype. CORBIDI 13725, adult male, collected on January 31, 2014 by L. Y. Echevarría.
Paratypes. Peru: Huancavelica Department: Tayacaja Province: CORBIDI 13637 a juvenile female from Jatuspata (12°15’14.617’’ S, 74°41’ 16.93’’ W) at 2,972 m asl, collected on September 3, 2013 by C. Landauro; CORBIDI 13638 a juvenile female from Pichiu (12°19’11.815’’ S, 74°39’3.588’’ W) at 2,036 m asl, collected on September 13, 2013 by C. Landauro; CORBIDI 13640 juvenile female from Pichiu, collected on September 12, 2013 by C. Landauro; CORBIDI 13641 adult female from Barropata (12°17’21.943’’ S, 74°41’19.035’’ W) at 1,725 m asl, collected on October 2, 2013 by C. Landauro; CORBIDI 13639 and CORBIDI 13642 female juveniles from Fundición (12°17’32.347’’ S, 74°39’45.433’’ W), at 2,169 m asl, collected on September 28, 2013 by C. Landauro; CORBIDI 13718 adult male from Chupto (12°19’52.96” S, 74°39’13” W) at 2,058 m asl, collected on January 23, 2014 by L. Y. Echevarría; CORBIDI 13719 adult male from Chupto (12°18’22.19” S, 74°39’18.42” W) at 2,314 m asl, on January 24, 2014 by L. Y. Echevarría; CORBIDI 13722 adult female from Chupto (12°19’52.96” S, 74°39’13” W) at 2,058 m asl, collected on January 21, 2014 by L. Y. Echevarría; CORBIDI 13728 adult female from Barropata (12°17’34.95” S, 74°41’34.08” W) at 1,704 m asl, collected on February 2, 2014 by L. Y. Echevarría; CORBIDI 13729 adult female from Barropata (12°17’22.11” S, 74°41’19.07” W) at 1,783 m asl, collected on February 3, 2014 by L. Y. Echevarría; CORBIDI 14669 adult female from Chupto (12°18’48.18” S, 74°39’1.83” W) at 2,345 m asl, collected on July, 1 2014 by L. Y. Echevarría; CORBIDI 14674 adult female from type locality, collected on July 6, 2014 by L. Y. Echevarría; CORBIDI 14676 juvenile female and CORBIDI 14677 adult male from Barropata (12°17’37.66” S, 74°41’39.18” W) at 1,693 m asl, collected on July 7, 2014 by L. Y. Echevarría; CORBIDI 15765–66 (adult male and female, respectively) and CORBIDI 15767 (juvenile male) from Pichiu (12°15’31.57” S, 75°6’3.6” W) at 2,206 m asl, collected on January 14, 2015 by L. Lujan; CORBIDI 16033 adult male from Pichiu (12°19’50” S, 74°39’13.07” W) at 2,090 m asl, collected on July 9, 2015 by J. Malqui; CORBIDI 20546–47 two adult males from San Luis de Estanque (12o19’8.62” S, 74o44’24.19” W) at 2,636 m asl., collected on Jun 17, 2019 by L.A. García-Ayachi. 
DiagnosisDiagnosis: Adult males of Stenocercus nigrobarbatus can be easily distinguished from all species of Stenocer- cus by having a continuous black patch covering the infralabials, throat, chest, ventral surfaces of forelimbs, belly (as a black midventral line), ventral surfaces of hind limbs, and pelvic region (Figs. 6B, 8B, 9B and 9C).
However, among the 69 currently described species of Stenocercus, S. nigrobarbatus resembles S. frittsi Tor- res-Carvajal, 2005, and S. variabilis Boulenger, 1901, by the following combinations of characters: (1) granular scales on the posterior surface of thighs, (2) imbricate and keeled lateral body scales, (3) a distinct row of enlarged vertebral scales, (4) unnotched gular scales, (5) three caudal whorls per autotomic segment, (6) brown dorsal ground color, and (7) distinct oblique neck fold, and antegular fold not continuous medially.
Stenocercus nigrobarbatus differs from the geographically close S. frittsi (character state of latter in parenthesis) by having a postfemoral mite pocket composed by one or more vertical folds or ridges, Type 1 of Torres-Carvajal (2007a) (postfemoral pocket absent in S. frittsi), 2–4 postrostrals (5–7), and two pairs of postxiphisternal inscrip- tional ribs, long, not in contact midventrally, pattern 1A of Torres-Carvajal (2004) (three pairs of postxiphisternal inscriptional ribs, two pairs long and the last pair short, not in contact midventrally; pattern 2B of Torres-Carvajal 2004).
Stenocercus nigrobarbatus can be distinguished from S. variabilis (character state in parenthesis) by having 2–4 postrostrals (6), postfemoral pocket composed by one or more vertical folds or ridges, Type 1 of Torres-Carva- jal (2007a) (slit-like opening, Type 2 of Torres-Carvajal 2007a), and inguinal groove absent (present). In addition, lateral body scales in S. nigrobarbatus are slightly smaller than dorsals, while S. frittsi and S. variabilis have lateral body scales approximately half the size of dorsals.


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Comment 
EtymologyThe specific epithet “nigrobarbatus” is a noun derived from the Latin words “nigro” (= black) and “barbatus” (= bearded). It refers to the black patch covering the gular region and ventral surface of neck, including the jaw angle, of adult males which resembles a black beard. 
References
  • VENEGAS, PABLO J.; LOURDES Y. ECHEVARRÍA, LUIS A. GARCÍA-AYACHI, CAROLL Z. LANDAURO 2020. Two new sympatric species of Stenocercus (Squamata: Iguania) from the inter-Andean valley of the Mantaro River, Peru. Zootaxa 4858 (4): 555–575 - get paper here
 
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