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Synophis niceforomariae PYRON, ARTEAGA, ECHEVARRÍA & TORRES-CARVAJAL, 2016

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Higher TaxaColubridae (Dipsadinae), Diaphorolepidini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Nicéforo María's Shadow Snake
S: Culebras Andinas de la Sombra de Nicéforo María 
SynonymSynophis niceforomariae PYRON, ARTEAGA, ECHEVARRÍA & TORRES-CARVAJAL 2016 
DistributionNC Colombia (Antioquia, Caldas, Tolima)

Type locality: Finca La Esperanza, Valle de La Manguita, Municipio Amalfi, Departamento Antioquia, Colombia (N6.978611, W-75.044444; 1394 m elevation).  
TypesHolotype: MHUA 14577 (Fig. 2), adult male collected 24 November 2007. Paratypes. MHUA14413 (Fig. 3), adult collected at Municipio Anori, Departamento Antioquia, Colombia (N6.987778, W-75.044444; 1320m); MLS2072, adult male collected at Municipio Campamento, Departamento Antioquia, Colombia (N6.979498, W-75.296602; 1656m). 
DiagnosisDiagnosis: Synophis niceforomariae can be differentiated from all other similar or related species by the combination of an unmodified vertebral scale row with a single weak keel; smooth nuchal scales; absence of a nuchal collar in adults; presence of a loreal; intranasals in contact; fused prefrontals; rostral concave; anal single; no apical pits; 2 postoculars; 10–12 infralabials; 8 supralabials; 184–193 ventrals; 127–131 subcaudals; and 19-19- 17 dorsal scale-rows. 
CommentSimilar species: This species was referred to as Synophis cf. bicolor by Pyron et al. (2015). 
EtymologyNamed after the French priest and naturalist Hermano Nicéforo María (born Antoine Rouhaire Siazaude), who established important natural-history collections at the Instituto de la Salle in Bogotá. While numerous taxa have been named mariarum, nicefori, or some derivative (see Adler 1989), he requested in a 1968 letter that he be addressed in print using his full name, Hermano Nicéforo María (Rodriguez 2002). Thus, it seems appropriate to use the specific epithet niceforomariae. 
  • PYRON, R. ALEXANDER; ALEJANDRO ARTEAGA, LOURDES Y. ECHEVARRÍA, OMAR TORRES-CARVAJAL 2016. A revision and key for the tribe Diaphorolepidini (Serpentes: Dipsadidae) and checklist for the genus Synophis. Zootaxa 4171 (2): 293–320 - get paper here
  • Sanchez TT, Montañez A, Bernal MH 2019. New records of Synophis lasallei (Nicéforo-María, 1950) and Synophis niceforomariae Pyron, Arteaga, Echevarría & Torres-Carvajal, 2016 (Serpentes, Colubridae) from the Cordillera Central in Colombia. Check List 15(6): 1051-1054 - get paper here
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