Tachymenoides goodallae LEHR, LUNDBERG, CUSI, SITES, TORRES & AGUILAR-PUNTRIANO, 2025
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Tachymenoides goodallae
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| Higher Taxa | Colubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) |
| Subspecies | |
| Common Names | E: Goodall’s Slender Snake |
| Synonym | Tachymenoides goodallae LEHR, LUNDBERG, CUSI, SITES, TORRES & AGUILAR-PUNTRIANO 2025: 7 Galvarinus tarmensis — TREVINE et al. 2022 Thamnodynastes sp. — LEHR et al. 2019 |
| Distribution | Peru (Pasco) Type locality: mountain ridge close to the radio tower at Chacos (10°39′05.7′′ S, 75°17′39.3′′ W; WGS84) in the buffer zone of the Yanachaga Chemill.n National Park, 2836 m a.s.l., Distrito Oxapampa, Provincia Oxapampa, Pasco, Peru |
| Reproduction | |
| Types | Holotype: MUSM 31152 (field number IWU 100), a male, collected on 15 February 2012 by E. Lehr and J. C. Cusi. Paratypes: n = 20 (4 females, 16 males), all from Peru, all collected by M. Lundberg. From along the road between Auquimarca and Uchuhuerta (10°43'29.2" S, 75° 41'57.3" W), 2700 m a.s.l., Distrito Paucartambo, Provincia Pasco, Pasco, collected on 09 February 2000 (MUSM 17813, ML 9, male), on 21 March 2000 (MUSM 17828, ML 15, male), and on 19 February 2001 (MUSM 18122, ML 122, male). From Auquimarca (10°44'54.2" S, 75°42'18.4" W), 2650 m a.s.l., Distrito Paucartambo, Provincia Pasco, Pasco, collected on 04 April 2000 (MUSM 17839, ML 22, female), collected on 21 March 2002 (MSUM 18580, ML 340, male), collected on 10 November 2002 (MUSM 19051, ML 456, male), and collected on 28 March 2004 (MUSM 23422, ML 680, female). From Huallamayo (10°45'53.2" S, 75° 43'25.1" W), 2500 m a.s.l., Distrito Paucartambo, Provincia Pasco, Pasco, collected on 13 March 2001 (MUSM 18104, ML 160, male). From Puagmaray (10°39'02.5" S, 75°46'35.5" W), 2600 m a.s.l., Distrito Huachon, Provincia Pasco, Pasco, collected on 26 April 2003 (MUSM 19167, ML 496, male), collected on 28 March 2003 (MUSM 19177, ML 484, male), collected on 29 April 2003 (MUSM 19187, ML 498, male), and collected on 23 March 2004 (MUSM 23420, ML 676, female). From the mountain ridge close to the radio tower at Chacos (10°39'30.1" S, 75◦17′52.2′′ W), 2780 m a.s.l., Distrito Oxapampa, Provincia Oxapampa, Pasco, collected on 06 November 2004 (MUSM 23485, ML 996, male; MUSM 23492, ML 1024, male), collected on 16 January 2005 (MUSM 24715, ML 1144, male; MUSM 24716, ML 1145, male), and collected on 24 March 2006 (MUSM 25403, ML 1400, male; MUSM 25422, ML 1399, male, GenBank accession number PV272666). From Ingenio (11◦27′01.2′′ S, 75◦41′10.6′′ W), 3050 m a.s.l., Distrito Tarma, Provincia Tarma, Jun.n, collected on 19 September 2004 (MUSM 23470, female [tissue sample MTD 2037], ML 917, GenBank accession numbers PV272665, PV255662). FromPacchani (14◦04′42.7′′ S, 69◦41′55.7′′ W), 2190m, Distrito Limbani, Provincia Sandia, Puno, collected on 06 November 2004 (MUSM23483,ML 987, male). |
| Diagnosis | Diagnosis: Usually 1 preocular, 10 maxillary teeth; 9 dentary teeth; dorsal scale rows smooth in 19/17/15 series; dorsal scales lacking apical pits; 140–157 ventrals in males (n = 17), 139–142 ventrals in females (n = 4); 52–67 paired subcaudals in males (n = 17), 52–56 paired subcaudals in females (n = 4); hemipenes with pronounced and deep spinulate calyces on the base of capitulum and distal portion of hemipenial body, extending onto half of the body on the asulcate side with the first proximal row of calyces distinctly larger, containing 9 calyces; in real life, the dorsum and flanks are olive brown to pale grayish brown with scattered black and cream flecks and no longitudinal stripes; the flanks are olive brown, dark brown, or dark gray, and darker than the dorsum toward the posterior body end; the head is laterally pale grayish brown or cream with a dark-brown lateral stripe from the nose to postoculars and diagonally to corner of the mouth, and contrasting dark-brown flecks on the cream supralabials and infralabials; the iris is brown with a distinct yellowish-tan ringlet; the ventral coloration is highly variable: nearly uniformly black, gray, and mottled dark gray; pale gray and mottled tan; or pale grayish tan; and usually, three irregularly shaped, narrow, longitudinal ventral stripes are present (one midventral and one ventrolateral on each side). (Lehr et al. 2025) Comparisons: see Appendix D, Table A3. Tachymenoides goodallae sp. nov. and Ta. affinis are similar in size (maximum SVL 433mmvs. 436mmin Ta. affinis, [1]) and have hemipenes with large calyces restricted to the hemipenial body on the asulcate side. However, both species can be distinguished as follows: Tachymenoides goodallae sp. nov. has dorsal scale rows mostly in 19/17/15 (61.9%, n = 13) series (17/17/15 in Ta. affinis, 81.3%, n = 13, [1]); wider hemipenial calyces (narrower, Appendix S S11 in [1] for CORBIDI 11791); cream supralabials with black blotches (black blotches absent); hemipenis with the first row of calyces distinctly larger (not distinctly larger); and a pale-gray venter with three continuous, longitudinal black stripes, one midventrally and one ventrolaterally on each side (two to five non-continuous longitudinal ventral stripes in a few specimens, forming a dark band in each ventral scale, with a general darkening tendency toward the tail; some specimens have an overall dark venter, with no discernible stripes [1]). Both Tachymenoides goodallae sp. nov. and Ta. harrisonfordi have dorsal scale rows in 19/17/15 series, supralabials with black blotches, but Ta. goodallae sp. nov. lacks longitudinal body folds (present in Ta. harrisonfordi), and the iris is brown (copper) with a distinct yellowish-tan ringlet (absent). Tachymenoides goodallae sp. nov., Ta. affinis, and Ta. harrisonfordi lack apical pits ([1,3], in this paper) and are immediately separated from all 15 Dryophylax spp., which have apical pits ([1,23]). Tachymenis currently contains four species: T. ocellata, T. peruviana, T. tarmensis, and T. trigonata. Tachymenoides goodallae sp. nov. has dorsal scales usually in 19/17/15 series whereas Tachymenis spp. has dorsal scales usually in 19/19/15 series [1]. Tachymenoides goodallae sp. nov. has contrasting dark-brown flecks on the cream supralabials and infralabials, whereas Tachymenis spp. has labials with a homogeneous clear background, with solid bands or small blotches under the eye [1]. Furthermore, the head of Ta. goodallae sp. nov. is shorter than the head of T. tarmensis (Figure 7). Galvarinus contains two species, G. chilensis (Schlegel, 1837 [24]) and G. attenuatus Walker, 1945 [5], which have dorsal scales in 19/19/15 series and apical pits (see Appendix S3 in [1]). The dorsal scales in Ta. goodallae sp. nov. are mostly in 19/17/15 rows (61.9%, n = 13), followed by 19/19/15 (19.0%, n = 4), 19/18/15 (14.3%, n = 3), and 19/17/14 (4.8%, n = 1); see Table A2. Trevine et al. [1] mention the dorsal scale row variation in Galvarinus as 19/19/15 (79%, n = 93), 19/19/17 (13%, n = 14), and one specimen in Tachymenis tarmensis as 19/19/13. Tachymenoides goodallae sp. nov. differs from G. attenuatus in having males with less subcaudals (52–67 [n = 17] vs. 69–75 in G. attenuatus, [1]) and females with less subcaudals (52–56 [n = 5] vs. 63–65 in G. attenuatus, [1]). (Lehr et al. 2025) Additional details (8461 characters) are available for collaborators and contributors. Please contact us for details. |
| Comment | |
| Etymology | Named after ethologist and conservationist Dr. Jane Goodall for her scientific accomplishments and worldwide conservation activism. |
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