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Toxicodryas pulverulenta (FISCHER, 1856)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Fischer's Cat Snake 
SynonymDipsas pulverulenta FISCHER 1856: 81
Dipsadomorphus pulverulentus — STERNFELD 1917
Dipsadomorphus Boueti CHABANAUD 1917: 373
Boiga pulverulenta — SCHMIDT 1923: 102
Boiga pulverulenta — MENZIES 1966
Toxicodryas pulverulenta — TRAPE & ROUX-ESTÈVE 1995: 40
Boiga pulverulenta — PITMAN 1974
Boiga pulverulenta — BROADLEY 1998
Toxicodryas pulverulenta — RÖDEL & MAHSBERG 2000
Boiga pulverulenta — HUGHES 2000
Toxicodryas pulverulenta — LEGRAND 2002
Boiga pulverulenta — BURGER et al. 2004
Toxicodryas pulverulenta — CHIRIO & LEBRETON 2007
Toxicodryas pulverulenta — ULLENBRUCH et al. 2010
Boiga pulverulenta — SEGNIAGBETO et al. 2012
Toxicodryas pulverulentus — WALLACH et al. 2014: 733
Toxicodryas pulverulentus — TRAPE & BALDÉ 2014
Boiga pulverulenta — CARLINO & PAUWELS 2015
Toxicodryas pulverulenta — SPAWLS et al. 2018: 512
Toxicodryas pulverulenta — GREENBAUM et al. 2021
Toxicodryas pulverulentus — BÖHME et al. 2023 
DistributionLiberia, Sierra Leone, Guinea (Conakry), Ivory Coast, Ghana, Togo, Benin, Nigeria, Gabon, Cameroon, Central African Republic, Democratic Republic of the Congo (Zaire), Congo (Brazzaville), Uganda, Angola, Republic of South Sudan (RSS) ?

Type locality: Edina, Grand Bassa County, Liberia (see comment)  
Reproductionoviparous 
TypesSyntype: ZMH R04376 (formerly no. 339), Formerly there were 2 syntypes. 
DiagnosisDiagnosis: A species of Toxicodryas restricted to West Africa, west of the Niger Delta, defined by the following combination of characters: maximum SVL < 1 meter (vs. maximum SVL > 1 meter in T. blandingii and T. vexator sp. nov.), DSRN 19–21 (vs. 23–25 in T. blandingii and 23–29 in T. vexator sp. nov.), DSRM 19–21 (vs. 21–25 in T. blandingii and T. vexator sp. nov.); cloacal plate undivided (vs. usually divided in T. blandingii, and divided or undivided in T. vexator sp. nov.); both sexes brown to pink with darker cross-bars that often enclose a whitish spot, and the dorsum and venter sprinkled with fine dark brown or black spots (vs. adult males glossy or velvety black with a yellow venter, and adult females light brown, gray, or yellowish-brown with light-brown or cream cross-bars on the flanks, with yellowish-brown venters in T. blandingii and T. vexator sp. nov.); hemipenis relatively long with long spines mid-way along the shaft that decrease in size towards the apex and base, with a domed apex (vs. hemipenis relatively short and massive [i.e., broad], proximal third covered with spines, distal two-thirds dimpled with a flattened apex in T. blandingii and T. vexator sp. nov.) (GREENBAUM et al. 2021).


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CommentDistribution: Has been reported from Sao Tome but records neen confirmation (SCHÄTTI & LOUMONT 1992); recent investigations show it actually doesn’t occur there (Ceríaco et al. 2018). The type locality has also been given as Sao Tomé [Is., West Africa], but that appears to be incorrect (Greenbaum et al. 2021,Loveridge 1958: 269).

Habitat: fully arboreal (Harrington et al. 2018). 
References
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  • Böhme, Wolfgang; Eberhard Fischer & Harald Hinkel 2023. Reptilien aus dem Irangi Forest, Provinz Kivu-Süd, Demokratische Republik Kongo. Sauria 45 (4): 29
  • Branch, W.R. & Rödel, M.-O. 2003. Herpetological survey of the Haute Dodo and Cavally forests, western ivory Coast, Part II: Trapping, results and reptiles. Salamandra 39 (1): 21-38 - get paper here
  • Briscoe, M.S. 1949. Notes on Snakes Collected in Liberia. Copeia 1949 (1): 16-18 - get paper here
  • Broadley, D.G. 1998. The reptilian fauna of the Democratic Republic of the Congo (Congo-Kinshasa). In: Schmidt, K.P. and Noble, G.K., Contributions to the Herpetology of the Belgian Congo... [reprint of the 1919 and 1923 papers]. SSAR Facsimile reprints in Herpetology, 780 pp.
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  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
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