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Trachylepis makolowodei CHIRIO, INEICH, SCHMITZ & LEBRETON, 2008

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Higher TaxaScincidae, Mabuyinae (Mabuyini), Scincoidea, Sauria, Squamata (lizards)
Common NamesE: Makolowodé’s Trachylepis 
SynonymTrachylepis makolowodei CHIRIO, INEICH, SCHMITZ & LEBRETON 2008
Trachylepis makolowodei — ALLEN et al. 2017 
DistributionCentral African Republic, Cameroon

Type locality: border of a forest river, near the village of Bélemboké, 45 km south of Nola in Central African Republic [about 3°17’ N, 16°15’ E]  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: MNHN-RA 1995.5612; see also Chirio & Ineich 2005), 31 May 1994, by Laurent Chirio; paratypes: MNHN-RA 1995.5610-5611, MNHN-RA 1995.5613-5614, MNHN-RA 1997.3464-3467, MNHN-RA 1997.3072-3073, MNHN-RA 1997.3075-3083 
DiagnosisDiagnosis: Moveable eyelids with a white border, the lower with a translucent palpebral disk; rectangular enlarged subocular, in direct contact with the lip and not reduced basally by the intrusion of adjacent supralabials; snout wide and massive; supra- and infralabials dark in formalin fixed specimens and mental plate often lighter; finger and toe lamellae not keeled and not spinose; absence of distinct longitudi- nal or transversal dorsal bands; back uniformly blackish and belly grey blue on formalin pre- served specimens and whitish on alcohol preserved specimens. This species can be distin- guished from all other members of the genus in western and central Africa (see Hoogmoed 194) by its elevated number of dorsal scale keels ( to 9), its size (up to 121 mm SVL), the relative length of its tail (approximately double the SVL), its high number of supraciliaries (6 to 8) and its coloration. It can be distinguished from T. polytropis, another sympatric species with an elevated number of dorsal keels, by the following combination of characters: large SVL, white marks on supralabials, black marks on supra- and infralabials which are weakly developed and most often absent, tail strongly flattened transversally, body only weakly flat- tened dorso-ventrally which gives the animal a massive habitus, presence of three well devel- oped ear lobules in front of tympanum, and finally the frequent occurrence of a large plate clearly individualised among the three largest plates located on the soles of the hand and foot. Its general scalation pattern is most similar to that encountered in T. boulengeri, but it can be distinguished from the later by its larger size (maximum known SVL121 mm vs. 90 mm of T. boulengeri), its head is proportionately larg- er, its higher number of supraciliaries and its coloration.
CommentSimilar species: This species has been confused with T. polytropis in the past. It is also similar to T. boulengeri.

Distribution: May also occur in the Democratic Republic of Congo. Not in Gabon fide Pauwels et al. 2018. 
EtymologyThe specific epithet makolowodei is a masculine patronym used in homage to Paul Makolowodé, originating from the village of Zimba, Central African Republic, who has worked as a field herpetologist in CAR and Cameroon (CAMHERP project). He has also collected the types of this species. 
  • Allen, Kaitlin E.; WALTER P. TAPONDJOU N., LUKE J. WELTON, AARON M. BAUER 2017. A new species of Trachylepis (Squamata: Scincidae) from Central Africa and a key to the Trachylepis of West and Central Africa. Zootaxa 4268 (2): 255–269 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Chirio, Laurent, Ivan Ineich, Andreas Schmitz and Matthew LeBreton. 2008. A new species of Trachylepis Fitzinger, 1843 (Squamata: Scincidae) from Central African forests. African Journal of Herpetology 57(1):13-28 - get paper here
  • Gvoždík, Václav; Matej Dolinay, Ange-G. Zassi-Boulou, Gabriel Badjedjea Babangenge 2018. New data on Trachylepis makolowodei from Central Africa. Herpetology Notes 11: 515-518 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
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