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Trachylepis punctulata (BOCAGE, 1872)

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Higher TaxaScincidae, Mabuyinae (Mabuyini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Speckled Skink, Speckled sand skink 
SynonymEuprepes punctulatus BOCAGE 1872: 76
Mabuia punctulata — BOULENGER 1887: 204
Mabuia punctulata — BOCAGE 1895: 44, 1897a: 195
Mabuya punctulata — SCHMIDT 1933: 12 [part]
Mabuya punctulata — FRADE 1963: 253
Mabuya punctulata — BROADLEY 2000: 100
Mabuya longiloba longiloba — LAURENT 1964: 73.
Mabuya variegata punctulata — BROADLEY 1974b: 7 [part]
Mabuya variegata punctulata — BRANCH 1998: 158
Mabuya damarana FITZSIMONS 1943: 224 (not PETERS) (part.)
Mabuya longiloba triebneri MERTENS 1954: 179
Mabuya variegata — BROADLEY 1971: 55
Mabuya variegata punctulata — BROADLEY 1975: 7
Mabuya punctulata — BROADLEY 2000: 100
Euprepis punctulatus — MAUSFELD et al. 2002
Trachylepis punctulata — BAUER 2003
Trachylepis punctulata — PORTIK & BAUER 2012
Trachylepis punctulata — LOBÓN-ROVIRA et al. 2022
Trachylepis punctulata — CERIACO et al. 2024: 99 
DistributionSW Angola, Namibia (excluding the coastal strip and the west, where replaced by M. variegata), W Zambia, botswana, N Cape, N and Free State Provinces of South Africa, NW Zimbabwe, Mozambique south of the Save River.

Type locality: Rio Coroca, dans le littoral au
sud de Mossamedes”, Namibe Province, Angola.

triebneri: Holotypus: SMF45559; male, Type locality: Osona bei Okahandja, Damaraland, SW-Afrika.  
Reproductionviviparous (not imputed, fide Zimin et al. 2022) 
TypesSyntypes: ZMB 6478, Rio Coroca, Angola, presented J.V. Barboza du Bocage; MBL 742 (2 specimens). The MBL syntypes were lost in the 1978 fire that destroyed the collection. See Bauer et al. (2003: 277) for discussion. 
DiagnosisDiagnosis: Ceriaco et al. 2024: 100 
CommentSynonymy: Mertens 1954 stated that his triebneri is a skink that had been called Mabuya damarana. However, his examination of the type of damaranus revelealed that it is identical with Mabuya varia Peters, hence he suggested to call Mabuya damarana Mabuya longiloba. Mertens distinguished two populations within longiloba, one of which he described as triebneri.

Distribution: see map in Ceriaco et al. 2024: 102 (Fig. 81). 
EtymologyPresumably named after the Latin punctula (small spot), plus the suffix -atus (having), in reference to the numerous small dark spots in this species.  
References
  • Bates, M. F. 1988. Life History Note: Mabuya variegata punctulata: Avian Predation. J. Herp. Assoc. Africa (34): 48-48 - get paper here
  • Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp.
  • Bocage, J. V. B. 1872. Diagnoses de quelques espéces nouvelles de Reptiles d'Afrique occidentale. Jorn. Sci. Math. Phys. Nat. Lisboa 4: 72-82. - get paper here
  • BOURQUIN, O. & V. GRUENER 2017. A NOTE ON THE HERPETOFAUNA OF MODISA CAMP (BOTSWANA) AND ITS SURROUNDINGS. African Herp News (64): 9-26 - get paper here
  • Broadley, D. G. 1971. The reptiles and amphibians of Zambia. The Puku, Occas. Pap. Dept. Wild. Fish. Natl. Parks Zambia 6: 1-143
  • Broadley, D. G. 1974. A review of the Mabuya lacertiformis complex in southern Africa. Arnoldia (Rhodesia) (ser. 7) 18: 1-16
  • Broadley, D.G. 2000. A review of the genus Mabuya in southeastern Africa (Sauria: Scincidae). African Journal of Herpetology, 49 (2): 87-110 - get paper here
  • Ceríaco, Luis M.P.; Mariana P. Marques, Diogo Parrinha, Arthur Tiutenko, Jeffrey L. Weinell, Brett O. Butler & Aaron M. Bauer 2024. The Trachylepis (Squamata: Scincidae) of Angola: An Integrative Taxonomic Review with the Description of Seven New Species Bulletin of the American Museum of Natural History Feb 2024 Vol. 2024, No. 465: 1-153 - get paper here
  • Conradie W, Keates C, Verburgt L, Baptista NL, Harvey J, Júlio T, Neef G. 2022. Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles. Amphibian & Reptile Conservation 16(2): 181–214 (e322) - get paper here
  • CONRADIE, W., DOUCETTE-RIISE, S., VANHOOYDONCK, B., ENGELBRECHT, H., MEASEY, G.J., & TOLLEY, K. 2011. Herpetological survey of Rooipoort Nature Reserve, Northern Cape, South Africa. African Herp News (53): 35-41 - get paper here
  • CONRADIE, W., ENGELBRECHT, H., HERREL, A., MEASEY, G.J., NIELSEN, S.V., V ANHOOYDONCK, B., & TOLLEY, K.A. 2011. Reptile survey of Venetia Limpopo Nature Reserve, Limpopo Province - South Africa. African Herp News (55): 6-12 - get paper here
  • FitzSimons, V.F. 1943. The lizards of South Africa. Transvaal Museum Memoir No.1 (Pretoria), 528 pp.
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Huey, R. B., Miles, D. B., & Pianka, E. R. 2021. Seasonality in Kgalagadi Lizards: Inferences from Legacy Data. American Naturalist 198 (6): 759-771 - get paper here
  • Lobón-Rovira J, Vaz Pinto P, S. Becker F, Tolley KA, Measey J, Bennet B, Boon B, de Sá S, Conradie W 2022. An updated herpetofaunal species inventory of Iona National Park in southwestern Angola. Check List 18(2): 289-321 - get paper here
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Mertens, R. 1954. Neue Eidechsen aus Südwest—Afrika. Senckenbergiana, 34:175—183
  • Pietersen, Darren W. Pietersen, Errol W. Haacke, Wulf D. 2013. First herpetological appraisal of the Parque Nacional de Banhine, Gaza Province, southern Mozambique. Annals of the Ditsong National Museum of Natural History 3: 153-163 - get paper here
  • Pietersen, Darren, Verburgt, Luke & Davies, John 2021. Snakes and other reptiles of Zambia and Malawi. Struik Nature / Penguin Random House South Africa, 376 pp., ISBN 9781775847373
  • Portik, Daniel M. & Aaron M. Bauer 2012. Untangling the complex: mole cular patterns in Trachylepis variegata and T. punctulata (Reptilia: Scincidae). African Journal of Herpetology 61 (2): 128-142; DOI:10.1080/21564574.2012.721808 - get paper here
  • SANTOS, B. S. .,MARQUES, M. P. ., BAUER, A. M. ., & CERÍACO, L. M. P. . 2021. Herpetological results of Francisco Newton’s Zoological Expedition to Angola (1903–1906): a taxonomic revision and new records of a forgotten collection. Zootaxa 5028 (1): 1–80 - get paper here
  • Tolley KA, Telford NS, Makhubo BG, Power RJ, Alexander GJ 2023. Filling the gap: Noteworthy herpetological discoveries in North West Province, South Africa. Zoosystematics and Evolution 99(1): 101-116 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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