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Trapelus agnetae (WERNER, 1929)

IUCN Red List - Trapelus agnetae - Least Concern, LC
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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common Names 
SynonymAgama agnetae WERNER 1929: 239
Agama pallidus haasi WERNER 1971
Trapelus pallidus agnetae — DISI et al. 2004
Trapelus pallidus agnetae — MODRY et al. 2004
Trapelus pallidus agnetae — SINDACO et al. 2006
Trapelus pallidus agnetae — SINDACO & JEREMČENKO 2008
Trapelus pallida haasi — AL-QURAN 2009
Trapelus agnetae — WAGNER et al. 2011
Trapelus pallidus haasi — AL-SHAMMARI 2012
Agama (Trapelus) pallidus hassi — AL-SADOON et al. 2016 (in error)
Trapelus pallidus haasi — MOHAMMED et al. 2017
Trapelus agnetae — BAR et al. 2021 
DistributionIsrael, Jordan, Iraq

Type locality: “Umgebung von Bir Molusi (Ka'ra) auf der Route der ehemaligen Wüstenpost, ungefähr halbwegs zwischen Damaskus und Bagdad.”

haasi: Jordan, Iraq; N Saudi Arabia, Kuwait; Terra typica: Azraq in Transjordan.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: NMW 23349 
DiagnosisDiagnosis: Small to medium sized member of the genus. Head longer than broad, limbs relatively long. Gular pouch absent. Ear opening longer than high, bordered by a row of spiny scales. Scales on dorsal surface of lower leg and basis of tail heterogeneous. Differs from T. pallidus in having a fringe of three to four distinct spines situated on the upper edge of the ear opening. Dorsal scales small, equal in size, forming a matrix with intermixed larger scales. Ventral scales usually smooth. 
CommentSynonymy: Wettstein 1951 thought that Agama agnetae is a juvenile Agama pallida.

Distribution: see Bar et al. 2021 for a map. 
EtymologyNamed after Ms. Agnes Gabriel, the collector of the type. 
References
  • Al-Quran, S. 2009. The Herpetofauna of the Southern Jordan. American-Eurasian J. Agric. & Environ. Sci., 6 (4): 385-391 [this journal has a dubious record, see http://goo.gl/81SBPv]
  • Al-Sadoon, Mohammed K., Bilal Ahmad Paray, Hamad S. Al-Otaibi 2016. Survey of the reptilian fauna of the Kingdom of Saudi Arabia. V. The lizard fauna of Turaif region. Saudi Journal of Biological Sciences, doi:10.1016/j.sjbs.2016.04.005 - get paper here
  • Al-Shammari, Ahmed M. 2012. Additional Records of Lizards in Ha'il Province, Saudi Arabia. Russ. J. Herpetol. 19 (4): 287-291 - get paper here
  • Aloufi, A., Amr, Z., & Baker, M. A. 2022. Reptiles from'Uruq Bani Ma'arid and Harat al Harrah protected areas in Saudi Arabia: Reptiles from two protected areas in Saudi Arabia. Herpetology Notes 15: 483-491
  • Aloufi, Abdulhadi A.; Zuhair S. Amr, Mohammad A. Abu Baker 2021. Reptiles and Amphibians of Al Madinah Al Munawwarah Province, Saudi Arabia. Russian Journal of Herpetology 28 (3): 1 - get paper here
  • Bar, A., Haimovitch, G. and Meiri, S. 2021. Field guide to the amphibians and reptiles of Israel. Edition Chimaira, Frankfurt Am Main. ISBN 9783-89973-120-0 - get paper here
  • Disi, Ahmad M., Zuhair S. Amr and Harald Martens 2004. On a collection of amphibians and reptiles made by J. Klapperich in Jordan. Herpetozoa 16 (3/4):141-150 - get paper here
  • Meiri, Shai; Amos Belmaker, Daniel Berkowic, Kesem Kazes, Erez Maza, Guy Bar-Oz and Roi Dor 2019. A checklist of Israeli land vertebrates. Israel Journal of Ecology and Evolution - get paper here
  • Modrý, P., Rifai, L., Abu Baker, M. & Amr, Z. 2004. Amphibians and reptiles of the Hashemite Kingdom of Jordan. Denisia 14 (Neue Serie 2): 407-420 - get paper here
  • Mohammed, Rihab Ghaleb; Fadhel Aabbas Rhadi, Nasrullah Rastegar-Pouyani, Eskandar Rastegar-Pouyani, Seyyed Saeed Hosseinian Yousefkhani 2017. Zoogeography of Lizards Fauna from Central and Southern Iraq with a Checklist of Iraqi Lizard’s Fauna. Russ. J. Herpetol. 24 (3): 193-201 - get paper here
  • Pola, L., Hejduk, V., Winkelhöfer, T., Zíka, A., Baker, M. A. A., & Amr, Z. S. 2020. Recent Observations on Amphibians and Reptiles in the Hashemite Kingdom of Jordan. Jordan Journal of Natural History, 7: 11-29 - get paper here
  • Scholz, S. et al. 2020. Fields Trughornviper (Pseudocerastes fieldi) - Beobachtungen im Lebensraum und bei der Fortpflanzung im Terrarium. Elaphe 2020 (3): 68-77
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Sindaco, R.; Serra, G. & Menegon, M. 2006. New data on the Syrian herpetofauna with a newly-recorded species of snake. Zoology of the Middle-East 37: 29-38 - get paper here
  • Sindaco, Roberto; Riccardo Nincheri, Benedetto Lanza 2014. Catalogue of Arabian reptiles in the collections of the “La Specola” Museum, Florence. Scripta Herpetologica. Studies on Amphibians and Reptiles in honour of Benedetto Lanza: pp. 137-164 - get paper here
  • WAGNER, PHILIPP; JANE MELVILLE, THOMAS M. WILMS and ANDREAS SCHMITZ 2011. Opening a box of cryptic taxa – the first review of the North African desert lizards in the Trapelus mutabilis Merrem, 1820 complex (Squamata: Agamidae) with descriptions of new taxa. Zoological Journal of the Linnean Society 163: 884–912 - get paper here
  • Werner, Franz 1929. Beitrage zur Kenntnis der Fauna von Syrien und Persien. Zool. Anz. 81 (7/10): 238-245 - get paper here
  • Werner, Y. L. 1971. Lizards and snakes from Transjordan, recently acquired by the British Museum (Natural Hisrory). Bulletin of the British Museum (Natural History), Zoology21: 213-256. - get paper here
  • Wettstein, O. von & Löffler, H. 1951. Ergebnisse der österreichischen Iran-Expedition 1949l50. Sitzungsber. Akad. Wiss. Wien, math.-nat. Kl., Abt. 1, 160: 427-448 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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