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Trimeresurus albolabris GRAY, 1842

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: White-lipped Tree Viper, White-lipped pitviper
G: Weisslippen-Bambusotter
Chinese: 白唇竹叶青蛇 
SynonymTrimesurus albolabris GRAY 1842: 48 (unjustified emendation)
Trimesurus albolabris — GRAY 1849
Lachesis gramineus albolabris — MELL 1922
Trimeresurus albolabris — POPE & POPE 1933
Trimeresurus albolabris albolabris — KRAMER 1977
Coluber gramineus — RAFFLES 1822 (nec Coluber gramineus SHAW 1802) (partim)
Trigonocephalus viridis — MÜLLER & SCHLEGEL 1842 (nec Vipera viridis DAUDIN 1803) (partim)
Trigonocephalus gramineus — CANTOR 1847 (nec Coluber gramineus SHAW 1802)
Trimeresurus gramineus — BOULENGER 1890 (nec Coluber gramineus SHAW 1802)
Lachesis gramineus — BOULENGER 1896 (nec Coluber gramineus SHAW 1802)
Lachesis grammineus (sic) — BRONGERSMA 1929
Bothrops erythrurus — LIDTH DE JEUDE 1890 (nec Trigonocephalus erythrurus CANTOR 1839)
Trimeresurus purpureomaculatus var. bicolor) — BOULENGER 1890 (nec Trimeresurus bicolor GRAY 1853, a synonym of Trigonocephalus erythrurus CANTOR 1839
Trimeresurus gramineus albolabris — MELL 1929
Trimeresurus albolabris — POPE & POPE 1933
Trimeresurus albolabris albolabris — WELCH 1994: 113
Trimeresurus albolabris — MANTHEY & GROSSMANN 1997: 407
Trimeresurus albolabris albolabris — COX et al. 1998: 20
Trimeresurus albolabris — MCDIARMID, CAMPBELL & TOURÉ 1999: 329
Cryptelytrops albolabris — MALHOTRA & THORPE 2004
Trimeresurus (Trimeresurus) albolabris — DAVID et al. 2011
Trimeresurus albolabris — WALLACH et al. 2014: 740
Cryptelytrops albolabris — CHAN-ARD et al. 2015: 284
Trimeresurus albolabris — WANG et al. 2022
Trimeresus albolabris — NEPALI & SINGH 2022 (in error)
Trimeresurus (Trimeresurus) albolabris— MIRZA et al. 2023 
DistributionNepal, Bhutan,
Burma, Thailand, Cambodia, Laos, Vietnam (Hoa Binh, Yen Bai, Hai Phong etc.),
S China (from Yunnan east to Guangdong and Hainan, north to Anhui; Fukien, Kwangsi, Fujian),
Indonesia (Sumatra, Java, Lombok, Sumbawa, Komodo, Sumba, Flores, Roti, Wetar, Kisar)

according to David & Vogel:

albolabris: Indonesia (Bangka, Sumatra, Java, Kalimantan [?], Madura, Sulawesi ?); Cambodia; Hong Kong; India (including Nicobar Islands); Laos; Macau; Myanmar; China (including Hainan); Thailand; Vietnam

Type locality: China  
TypesHolotype: BMNH 1946.1.19.85 
DiagnosisDIAGNOSIS (DIAGNOSTIC CHARACTERS). Scales in 21 (rarely 19) longitudinal rows at midbody; 10–11(12) upper labials, the first partially or completely fused to the nasal; head scales small, subequal, feebly imbricate, smooth or weakly keeled; supraoculars narrow, occasionally enlarged, undivided, 8–12 interocular scales between them; temporal scales smooth; green above, side of head below eyes yellow, white or pale green, much lighter than rest of head; below, green, yellowish or white below, a light ventrolateral stripe present in all males, absent in females; end of tail not mottled brown; ventrals: males 155–166, females 152–176; subcaudals: males 60–72, females 49–66, paired; hemipenes without spines. Total length males 600 mm, females 810; tail length males 120 mm, females 130 mm. [after LEVITON 2003] 

Distribution: Not in India after Vogel et al. 2022 restricted its range to other regions. Specimens from India (Nagpur) have been re-identified as T. salazar (Vogel et al. 2022). Has been reported from the Andaman and Nicobar Islands but no voucher specimens exist according to DAS 1999. Specimens from Nicobar have been described as a new species, Trimeresurus davidi Chandramouli et al. 2020. See map in Vogel et al. 2023: 529 (Fig. 5) for localities.

Subspecies: The subspecies insularis and septentrionalis have been raised to full species status by GIANNASI et al. (2001) based on RFLPs (which may or may not be reliable markers for (sub-) species boundaries).

Distribution: Possibly in Bhutan (Lenz 2012). Records from Sulawesi are questionable (DE LANG & VOGEL 2006). Not on Timor (H. Kaiser, pers. comm.).

Habitat: fully arboreal (Harrington et al. 2018). 
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