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Tropidodipsas papavericola GRÜNWALD, TORIBIO-JIMÉNEZ, MONTAÑO-RUVALCABA, FRANZ-CHÁVEZ, PEÑALOZA-MONTAÑO, BARRERA-NAVA, JONES, RODRIGUEZ, HUGHES, STRICKLAND & REYES-VELASCO, 2021

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Poppyfield Snailsucker
S: Caracolera amapolera 
SynonymTropidodipsas papavericola GRÜNWALD, TORIBIO-JIMÉNEZ, MONTAÑO-RUVALCABA, FRANZ-CHÁVEZ, PEÑALOZA-MONTAÑO, BARRERA-NAVA, JONES, RODRIGUEZ, HUGHES, STRICKLAND & REYES-VELASCO 2021: 242 
DistributionMexico (Guerrero)

Type locality: Mexico: Guerrero, 12.5 km S of Puerto del Gallo on road from Nuevo Dehli to Puerto del Gallo, Municipio de Atoyac de Álvarez, 1914 m a.s.l.  
Reproduction 
TypesHolotype: INIRENA 2801, Adult male. SVL 421 mm; TL 137 mm; TotL 558 mm. field number CIG 1495.
Paratypes: (n=5) INIRENA 2802 (original field number CIG 1496, Fig. 5C). Sub-adult male, collected AOR 18.1 km S of Puerto del Gallo on road from Nuevo Dehli to Puerto del Gallo, Municipio de Atoyac de Álvarez (17.4342°, -100.191°, datum=WGS84, 1654 m a.s.l.), Guerrero, Mexico on June 29th, 2019 by Christoph I. Grünwald, Miguel A. Peñaloza-Montaño, Eduardo Y. Barrera-Nava, Karen I. Morales-Flores and Janelle Morales-Flores. INIRENA 2803 (original field number CIG 1502). Sub-adult male collected DOR at 5 km S of La Laguna, on road from San Luis La Loma to Bajitos de la Laguna, Municipio de Técpan de Galeana (17.5137°, -100.7737°, datum=WGS84, 1686 m a.s.l.), Guerrero, Mexico on June 29th, 2019 by Jason M. Jones and Carlos Montaño-Ruvalcaba. INIRENA 2804 (original field number CIG 1632, Figs 5A–B). Adult male, collected at Bajitos de la Laguna, Municipio de Técpan de Galeana, Guerrero, Mexico on October 30th, 2019 and retrieved by Alejandro Lara and donated to Jason M. Jones. INIRENA 2805 (original field number CIG 1457). Sub-adult male, collected at Jaguar Research Facility, Municipio de Técpan de Galeana, Guerrero, Mexico on June 10th, 2019 by Miguel A. Peñaloza-Montaño and Eduardo Y. Barrera-Nava. INIRENA 2810 (JRV 0362, Fig. 2D). Juvenile male, collected DOR at 4.2 km S of La Laguna, on San Luis San Pedro - La Laguna Rd., Municipio de Técpan de Galeana (17.5173°, -100.7715°, datum=WGS84, 1620 m a.s.l.), Guerrero, Mexico on July 17th, 2020 by Jason M. Jones 
DiagnosisDiagnosis Tropidodipsas papavericola sp. nov. is placed in the genus Tropidodipsas based on phylogenetic evidence (Fig. 6). It belongs to the Tropidodipsas fasciata group as defined by Kofron (1987) based on possessing a laterally compressed body, head distinctly wider than neck, protruding eyes of moderate to large diameter, vertebral and paravertebral scales not wider than other dorsal scales, 17 maxillary teeth, 22 dentary teeth, postmental scale absent. The species differs from all described species of Tropidodipsas recognized herein based on the following combination of characters: (1) 15/15/15 rows of smooth dorsal scales with no enlarged vertebral row; (2) prefrontal broader than long, not entering orbit; (3) loreal almost square, slightly longer than broad, not entering orbit; (4) 179–189 ventral scales in males, unknown in females; (5) 69–76 divided subcaudals in males, unknown in females; (6) 25–34 pale bands on body and 10–16 pale bands on tail; (7) 26–36 dark bands on body and 10–16 on tail, irregular with faded pale centers in adult individuals; (8) TL/SVL 0.30–0.35 in males, unknown in females; (9) iris dark chocolate-gray in life. (Grünwald et al. 2021) 
Comment 
EtymologyNamed after papaver + cola, referring to the living among poppy plants (genus Papaver) which are illegally planted throughout the range of this species for the extraction of opium gum. 
References
  • Grünwald CI, Toribio-Jiménez S, Montaño-Ruvalcaba C, Franz-Chávez H, Peñaloza-Montaño MA, Barrera-Nava EY, Jones JM, Rodriguez CM, Hughes IM, Strickland JL, Reyes-Velasco J 2021. Two new species of snail-eating snakes of the genus Tropidodipsas (Serpentes, Dipsadidae) from southern Mexico, with notes on related species. Herpetozoa 34: 233-257 - get paper here
 
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