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Tympanocryptis diabolicus DOUGHTY, KEALLEY, SHOO & MELVILLE, 2015

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Hamersley pebble-mimic dragon 
SynonymTympanocryptis diabolicus DOUGHTY, KEALLEY, SHOO & MELVILLE 2015
Tympanocryptis diabolicus — CHAPPLE et al. 2019: 93 
DistributionAustralia (Western Australia: Hamersley Range in the Pilbara region)

Type locality: Mount Brockman (22°17'31"S, 117°16'23"E), Western Australia  
Reproduction 
TypesHolotype: WAM R135413, an adult male collected on 20 November 1998 by S. Anstee (Fig. 10). Paratypes. WAM R135411 (male), Mount Brockman (22°17'31"S, 117°16'23"E); WAM R135412 (male), Mount Brockman (22°17'31"S, 117°16'23"E), WAM R135454 (male), Mount Brockman (22°17'31"S, 117°16'23"E), WAM R170204 (male), 53 km north-north-west of Tom Price (22°18'21"S, 117°28'47"E); WAM R170281 (male), 51 km east-south-east of Paraburdoo (23°17'33"S, 118°09'23"E) 
DiagnosisDiagnosis: Distinguished from other Tympanocryptis by the following combination of character states: presence of two pre-cloacal pores, lack of longitudinal stripes on the dorsum, presence of enlarged scales with raised spines arranged in transverse rows of 2–5 scales or scattered on dorsum, snout straight or concave, scales on snout smooth to rugose with feeble keels, rostral width ~2–3 times height, keels on scales of upper arm aligned, well-defined row of enlarged scales at anterior and dorsal edge of thigh forming conspicuous ridge, scales on dorsal surface of thigh aligned, ventrals with low keels, and rich reddish-brown ground coloration with some irregular dark blotching along midline of dorsum. 
Comment 
Etymology‘diabolicus’ means ‘devil’ in Latin, in reference to the rich red coloration possessed by most individuals of this species. Used as a noun in apposition. 
References
  • Chapple, David G.; Reid Tingley, Nicola J. Mitchell, Stewart L. Macdonald, J. Scott Keogh, Glenn M. Shea, Philip Bowles, Neil A. Cox, John C. Z. Woinarski 2019. The Action Plan for Australian Lizards and Snakes 2017. CSIRO, 663 pp. DOI: 10.1071/9781486309474 - get paper here
  • DOUGHTY, PAUL; LUKE KEALLEY, LUKE P. SHOO & JANE MELVILLE 2015. Revision of the Western Australian pebble-mimic dragon species-group (Tympanocryptis cephalus: Reptilia: Agamidae). Zootaxa 4039 (1): 085–117 - get paper here
  • Ellis, Ryan J. 2019. An annotated type catalogue of the dragon lizards (Reptilia: Squamata: Agamidae) in the collection of the Western Australian Museum. Records of the Western Australian Museum 34: 115–132 - get paper here
 
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