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Tympanocryptis macra STORR, 1982

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Savannah Earless Dragon (also as Savanna Earless Dragon) 
SynonymTympanocryptis lineata macra STORR 1982: 61
Tympanocryptis lineata macra — SHOO et al. 2008
Tympanocryptis lineata macra — COGGER 2014: 758
Tympanocryptis lineata macra — MELVILLE et al. 2019
Tympanocryptis lineata macra — ELLIS 2019
Tympanocryptis macra — MELVILLE et al. 2019 
DistributionAustralia (N Western Australia, N Northern Territory)

Type locality: 16 km S of main dam at Lake Argyle, Western Australia, in 16°15'S, 128°40'E.  
TypesHolotype: WAM R44553 
DiagnosisDiagnosis (macra): A moderately large, relatively slender subspecies of T. lineata Peters, most like T. l. centralis Sternfeld but larger and having longer limbs and tail and more subdigital lamellae.

Diagnosis. A species of Tympanocryptis with rostral scale continuous with the canthus rostralis, no lateral neck fold, keels of dorsal snout and head scales all aligned parallel to the midline, weak five- lined colour pattern, keeled ventrals and throat scales, and large size (Melville et al. 2019: 29).

Comparison with other species. Tympanocryptis macra is a distinctive species, the largest species in the genus and the most gracile, with relatively longer extremities [21]. Its weakly contrasting colour pattern, as Storr [21] noted, makes it superficially similar to T. rustica, but its tapered snout and cephalic keels oriented in parallel series immediately distinguish T. macra from either T. centralis or T. rustica. Recent publications (e.g. [45]) have at times confused this species with T. uniformis, a species that is currently poorly understood, but markedly distinct from T. macra in its round-headed, ‘dumpy’ habitus and dorsal surface lacking enlarged spinous scales (Melville et al. 2019: 29). 
CommentSubspecies: Tympanocryptis lineata macra has been elevated to full species status.

Distribution: see map in Melville et al. 2019: 6 (Fig. 1)

See also Tympanocryptis lineata.

Habitat. A wide range of tropical savannah grasslands and shrublands, mainly on cracking soils. Males observed perching on low rocks or clumps of soil using tripod stance, similar to that observed previously in T. tetraporophora. Will flee down cracks when alarmed. Has also been observed perched off the ground in grasses during the heat of the day. 
EtymologyThe name macra is Latin for 'lean'. 
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