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Tympanocryptis tetraporophora LUCAS & FROST, 1895

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards) 
Subspecies 
Common NamesEyrean earless dragon, Long-tailed Earless Dragon 
SynonymTympanocryptis tetraporophora LUCAS & FROST 1895
Tympanocryptis lineata tetraporophora — MITCHELL 1948: 72
Tympanocryptis tetraporophora — WERMUTH 1967: 99
Tympanocryptis tetraporophora — SMITH et al. 1999
Tympanocryptis tetraporophora — COGGER 2000: 356
Tympanocryptis tetraporophora — WILSON & SWAN 2010 
DistributionNE W Australia (Kimberley-Division)
Australia (New South Wales, Northern Territory, Queensland, South Australia, Victoria)

Type locality: Adminga and Dalhousie, South Australia.  
Reproductionoviparous 
TypesLectotype: NMV D7701, male, from either Adminga or Dalhousie, S. A., designated by Coventry, A. J. (1970). 
DiagnosisDiagnosis. A medium-sized Tympanocryptis with very variable body patterning, consisting of darker brown broken cross- bands on body. A weak to strong single pale ventral stripe. Dorsal scales homogenous with scattered enlarged mucronate scales, enlarged scales not substantially larger than other scales. Dorsal head scales homogenous, weakly-strongly keeled with scattered enlarged mucronate scales mainly to rear of head and neck. Tail scales strongly keeled and tail tapers to rounded tip. Dorsal patterning variable, 4–6 broken darker cross-bands on torso on a reddy-brown or brown background. Dark brown bands on legs and also extending down 2/3rds of the tail length. Banding does not extend to ventral surfaces. Ventral scales weakly to strongly keeled. Some individuals have dark pigmentation on ventral surface of head, throat and upper chest. Nostril nearer eye than tip of snout. Four pores present, two pre-anal and two femoral pores (from Melville et al. 2014).
 
CommentT. tetraporophora can be distinguished from T. cephalus, T. lineata, and T. intima by the presence of four pores (two preanal and two femoral). T. tetraporophora can be distinguished from T. pentalineata sp. nov by lacking five longitudinal body stripes.

Distribution: The distribution of T. tetraporophora overlaps with a number of other Tympanocryptis, including T. cephalus, T. lineata, T. intima and T. pentalineata sp. nov. 
EtymologyNamed for the presence of four pores, two pre-anal and two femoral. 
References
  • Ackermann, T. 2006. Nutzung eines Wintergartens als Terrarienstandort für ein Wüstenterrarium. Teil 2: Tympanocryptis tetraporophora. Reptilia (Münster) 11 (60): 62-67 - get paper here
  • Ackermann, T. & Fritz, P. 2006. Australische Taubagamen Tympanocryptis tetraporophora. Iguana Rundschreiben 19 (1): 17-28
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp.
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Coventry, A. J. 1970. Reptile and amphibian type specimens housed in the National Museum of Victoria. Mem. Natl. Mus. Vic. 31: 115-124.
  • Freynik, C. & Drewes, O. 2011. Die Bartagame, Zwergbartagame & Australische Taubagame. Vivaria-Verlag, Meckenheim, 80 pp. [review in Reptilia #98: 82]
  • Fritz, P. & Ackermann, T. 2012. Die Australische Taubagame, Tympanocryptis tetraporophora. Natur und Tier Verlag, 64 pp. [review in Iguana 25 (2): 21] - get paper here
  • Greenbaum, E. 2000. Herpetofaunal observations in disparate habitats in south Sustralia, New South Wales, and queensland, Australia. Herpetological Bulletin (72): 6-16 - get paper here
  • Greer, A.E. and S. Smith 1999. Aspects of the morphology and reproductive biology of the Australian earless dragon lizard Tympanocryptis tetraporophora. Australian Zoologist 31(1):1-16
  • Lucas, A. H. S., and C. Frost. 1895. Preliminary notice of certain new species of lizards from central Australia. Proceedings of the Royal Society of Victoria, 7: 264-269 - get paper here
  • Melville J, Smith K, Hobson R, Hunjan S, Shoo L 2014. The Role of Integrative Taxonomy in the Conservation Management of Cryptic Species: The Taxonomic Status of Endangered Earless Dragons (Agamidae: Tympanocryptis) in the Grasslands of Queensland, Australia. PLoS One 9 (7): e101847. doi:10.1371/journal.pone.0101847 - get paper here
  • Mitchell, F. J. 1948. A revision of the lacertilian genus Tympanocryptis. Rec. South Austral. Mus. 9: 57-86 - get paper here
  • Müller, Hans D. 1998. Die australische Agame Tympanocryptis tetraporophora im Terrarium. Elaphe 6 (4): 2-6
  • P., Simone 2016. Professionelle Nachzucht von Terrarientieren für den Zoofachhandel Reptilia 21 (118): 52-55 - get paper here
  • Smith WJS, Osborne WS, Donnellan SC & Cooper PD 1999. The systematic status of earless dragon lizards, Tympanocryptis (Reptilia: Agamidae), in south-eastern Australia. Australian Journal of Zoology 47: 551-564 - get paper here
  • Storr, G M 1982. Taxonomic notes on the genus Tympanocryptis Peters (Lacertilia: Agamidae). Rec. West. Austr. Mus. 10 (1): 61-66 - get paper here
  • Storr, G. M. 1964. The agamid lizards of the genus Tympanocryptis in Western Australia. J R. Soc. West. Aust. 47: 43-50.
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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