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Tympanocryptis tolleyi MELVILLE, CHAPLIN, HIPSLEY, SARRE, SUMNER & HUTCHINSON, 2019

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards) 
Subspecies 
Common NamesE: Gawler Earless Dragon 
SynonymTympanocryptis tolleyi MELVILLE, CHAPLIN, HIPSLEY, SARRE, SUMNER & HUTCHINSON 2019: 24 
DistributionAustralia (South Australia)

Type locality: 2.8 km ESE Malbooma Outstation, Mulgathing Station, South Australia; 30°41’4700 S, 134°12’2100 E.  
Reproduction 
TypesHolotype. SAMA R69147; adult male. Collected by M.H. on 13 October 2015.
Paratypes. SAMA R26711, 30 km SW Mabel Ck H/S, South Australia, 29°100 3000 S, 134°080 E, adult female;
R39978-79, 2 km SSW of Grace Bore, Bulgunnia Station, South Australia, adult and juvenile females; R44704, Mt Vivian Yards, The Twins Station, South Australia, 30°36’ S, 135°42’ E, adult female; R45143, Nth Halls Paddock, Wilgena Station, South Australia, 30°3500300 S, 134°1903700 E, adult male; R47558, 1.9 km NE of Crows Nest Bore, Mt Vivian Station, South Australia, 30°49’2200 S, 135°49’1800 E, adult male; R58048, 25 km NE Half Moon Lake, South Australia, 29°52’3400 S, 133°37’0500 E, adult male; R62565, Challenger Mine lease, 23.6 km SE Blowout Trig, South Australia, 29°52’3200 S, 133°34’5000 E, adult female; R65637, 8.4 km NE Bon Bon Homestead, South Australia, 30°21’4900 S, 135°31’4500 E, adult male; R65808, 5.12 km NE Mullaquana Homestead, 33°10’3800 S, 137°24’0800 E, adult male; R69085, 0.2 km SSW Malbooma Outstation, Mulgathing Station, South Australia, 30°41’1700 S, 134°10’4000 E, adult female; R69141, 1.1 km SE Malbooma Outstation, Mulgathing Station, South Australia, 30°41’3800 S, 134°11’1000 E, adult male. 
DiagnosisDiagnosis. A species of Tympanocryptis with rostral scale separated from the canthus rostralis, nasal scale extending dorsally across the canthus and bordered below by enlarged scales, well-developed lateral neck fold, keeled dorsal head scales, strong sexual dimorphism in dorsal pattern (cross-banded females, males with longitudinal light dorsal lines that coalesce medially), ventral surface white. 
CommentHabitat. Arid chenopod shrublands and grasslands and the skirts of stony hills and rock outcrops, generally with sparse overstorey and with gravelly or heavier soil types. In the vicinity of the type locality at Malbooma outstation and the Kychering Rocks, lizards inhabit both the sparsely vegetated granitic and sedimentary rock outcrops and the surrounding gravel and clay plains.

Distribution: see map in Melville et al. 2019: 6 (Fig. 1: species D) 
EtymologyNamed for Timothy David Tolley of the South Australian Museum Waterhouse Club, in recognition of his long-term support for field research by the South Australian Museum, including coordination of the Waterhouse Club expedition that yielded the series of specimens that includes the holotype of this species. 
References
  • Melville J, Chaplin K, Hipsley CA, Sarre SD, Sumner J, Hutchinson M. 2019. Integrating phylogeography and high-resolution X-ray CT reveals five new cryptic species and multiple hybrid zones among Australian earless dragons. R. Soc. open sci. 6: 191166 - get paper here
 
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