Tytthoscincus aesculeticola (INGER, LIAN, LAKIM & YAMBUN, 2001)

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Higher Taxa	Scincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies
Common Names
Synonym	Sphenomorphus aesculeticola INGER, LIAN, LAKIM & YAMBUN 2001 Sphenomorphus aesculeticola — LINKEM et al. 2010 Tytthoscincus aesculeticola — LINKEM et al. 2011
Distribution	Malaysia (Borneo: Sabah) Type locality: Mount Lamaku (1350 m) (4° 52’ N, 115° 38’ E), 14,6 km from Mendolong, Sipitang District, Sabah, Malaysia.
Reproduction	oviparous (not imputed, fide Zimin et al. 2022)
Types	Holotype: FMNH 239852
Diagnosis	Diagnosis: A short-legged, small (SVL<45 mm) species of Sphenomorphus having four supraoculars, fewer than 10 lamellae under the fourth toe; scale rows usually 28-30, and prefrontals not meeting in the midline (Inger et al. 2001: 182). Additional details (2087 characters) are available for collaborators and contributors. Please contact us for details.
Comment
Etymology	Named after Latin “aesculetum” = oak forest and “cola” = dweller.
References	Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo Grismer, L. Lee 2006. Two new species of skinks (Genus Sphenomorphus Fitzinger 1843) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History 9 (2): 151-162. Grismer, L. Lee. 2008. A new species of insular skink (Genus Sphenomorphus Fitzinger 1843) from the Langkawi Archipelago, Kedah, West Malaysia with the first report of the herpetofauna of Pulau Singa Besar and an updated checklist of the herpetofauna of Pulau Langkawi. Zootaxa 1691: 53-66 - get paper here GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species. Zootaxa 4092 (2): 231–242 - get paper here Grismer, L.L. 2007. A new species of small montane forest floor skink (genus Sphenomorphus Fitzinger 1843) form Southern peninsular Malaysia. Herpetologica 63 (4): 544-551 - get paper here Grismer, L.L.; Ahmad, N. & Kin Onn, C. 2009. A new, diminutive, upland Sphenomorphus Fitzinger 1843 (Squamata: Scincidae) from the Belum-Temengor Forest Complex, Peninsular Malaysia. Zootaxa 2312: 27–38 - get paper here Grismer, L.L.; Wood Jr., P.L. & Grismer, J.L. 2009. A New Insular Species of Skink of the Genus Sphenomorphus Strauch 1887 (Squamata: Scincidae) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. Tropical Life Sciences Research, 20(1): 51–69 - get paper here Inger, R.F.; Lian, R.F.; Lakim, M. & Yambun, P. 2001. New species of the lizard genus Sphenomorphus (Lacertilia: Scincidae), with notes on ecological and geographic distribution of species in Sabah, Malaysia. Raffles Bull. Zool. 49 (2): 181-189 - get paper here Linkem, C.W.; Diesmos, A.C.; Brown, R.M. 2010. A NEW SPECIES OF SCINCID LIZARD (GENUS SPHENOMORPHUS) FROM PALAWAN ISLAND, PHILIPPINES. Herpetologica 66 (1): 67–79 - get paper here Linkem, Charles W.; Arvin C. Diesmos, Rafe M. Brown 2011. Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163: 1217–1243 - get paper here Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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