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Tytthoscincus atrigularis (STEJNEGER, 1908)

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Zamboanga Sphenomorphus 
SynonymSphenomorphus atrigularis STEJNEGER 1908: 199
Sphenomorphus atrigularis — TAYLOR 1922
Lygosoma (Sphenomorphus) atrigularis — SMITH 1937: 221
Sphenomorphus atrigularis — MITTLEMAN 1952: 22
Sphenomorphus atrigularis — BROWN & ALCALA 1980: 158
Tytthoscincus atrigularis — LINKEM et al. 2011
Tytthoscincus atrigularis — GRISMER et al. 2016 
DistributionPhilippines (Mindanao, Zamboanga Peninsula, Basilan)

Type locality: Mindanao Island.  
Reproductionoviparous (phylogenetic imputation, fide Zimin et al. 2022) 
TypesHolotype: USNM 37386, Datu Ahib's place, base of Malingdang Mtns, Catagan, Misamis Province, Mondanao, Philippines, collected E.A. Mearns, 24.v.1906. 
DiagnosisDiagnosis: No supranasals; fronto-nasal broadly in contact with rostral, fronto-parietals distinct; four large supraoculars; no auricular lobules; distance between tip of snout and fore leg about once and a half in distance between axil and groin; fourth toe scarcely longer than third; head and body nearly equalling length of tail; temporal region covered by scales scarcely larger than dorsals [from STEJNEGER 1908]. 
Comment 
References
  • Brown, R.M. et al. 2010. Species boundaries in Philippine montane forest skinks (Genus Sphenomorphus): three new species from the mountains of Luzon and clarification of the status of the poorly known S. beyeri, S. knollmanae, and S. laterimaculatus. Scient. Pap. Nat. Hist. Mus. Univ. Kansas (42): 1-27
  • Brown, Rafe M., Jimmy A. McGuire, John W. Ferner and Angel C. Alcala 1999. New species of diminutive scincid lizard (Squamata: Lygosominae: Sphenomorphus) from Luzon Island, Republic of the Philippines. Copeia 1999 (2): 362-370 - get paper here
  • Brown, WC. & A.C. ALCALA 1980. Philippine Lizards of the family Scincidae. Silliman Univ. Nat. Sci., Dumaguete City, Mon., Ser. 2: i-xi + 1-246.
  • GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species. Zootaxa 4092 (2): 231–242 - get paper here
  • Linkem, Charles W.; Arvin C. Diesmos, Rafe M. Brown 2011. Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163: 1217–1243 - get paper here
  • Mittleman, M.B. 1952. A generic synopsis of the Lizards of the Subfamily Lygosominae. Smithson. misc. Collns, 117: 1-35. - get paper here
  • Smith, M.A. 1937. A review of the genus Lygosoma (Scincidae: Reptilia) and its allies. Records of the Indian Museum 39 (3): 213-234
  • Stejneger,L. 1908. Three new species of lizards from the Philippine Islands. Proc. US Natl. Mus. 34: 199-204 - get paper here
  • Taylor, E.H. 1922. The lizards of the Philippine Islands. Department of Agriculture and Natural Resources, Bureau of Science, Government of the Philippine Islands, Manila, Publication no. 17: 269 pp. - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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