Tytthoscincus hallieri (LIDTH DE JEUDE, 1905)

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Higher Taxa	Scincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies
Common Names
Synonym	Lygosoma hallieri LIDTH DE JEUDE 1905: 197 Lygosoma hallieri — DE ROOIJ 1915: 210 Lygosoma (Sphenomorphus) hallieri — BRONGERSMA 1942: 129 Sphenomorphus hallieri — BACON 1967: 72 Sphenomorphus hallieri — INGER et al. 2001 Sphenomorphus hallieri — LINKEM et al. 2010 Tytthoscincus hallieri — LINKEM et al. 2011 Tytthoscincus hallieri — GRISMER et al. 2016
Distribution	Indonesia (Borneo), Malaysia (Sabah) Type locality: Putus Sibau, Kalimantan Barat (Borneo), Indonesia.
Reproduction	oviparous (not imputed, fide Zimin et al. 2022)
Types	Holotype: RMNH (Leiden) 4456
Diagnosis	Diagnosis (species). Prefrontals juxtaposed; parietals small, separated from supraoculars; tympanum slightly sunken, without auricular lobules; 4 supraoculars; 36-39 scale rows; limbs moderately developed, hind limbs longer than distance between anterior corner of eye and the forelimb; 2 pairs of chin shields (from BACON 1967).
Comment	Habitat: terrestrial. Type species: Lygosoma hallieri LIDTH DE JEUDE 1905 is the type species of the genus Tytthoscincus LINKEM, DIESMOS & BROWN 2011. Description (genus): Tytthoscincus can be identified by the following characters: (1) body size diminutive, usually less than 45 mm SVL; (2) temporal scales small, same size and shape as lateral body scales (Fig. 5); and (3) digits small, toe IV slightly longer than, or equal to, toe III.
Etymology	Named after Dr. Johann Gottfried "Hans" Hallier (1868-1932) a botanist who studied in Munich and Jena and later worked at the universities of Göttingen, Munich, Hamburg, and Leiden. Hewas on the first Dutch scientific expedition to Borneo (1893-1894) and traveled and collected in the Philippines (1903-1904).
References	Bacon, J. P. 1967. Systematic status of three scincid lizards (genus Sphenomorphus) from Borneo. Fieldiana: Zoology 51: 63-76. - get paper here Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Brongersma,L.D. 1942. Notes on scincid lizards. Zoologische Mededelingen 24: 125-152 - get paper here Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo Das, I. & Yaakob, N. 2007. Status of knowledge of the Malaysian herpetofauna. In Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. in: L. S. L. Chua, L. G. Kirton & L. G. Saw (eds.), Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31-81 de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here Grismer, L. Lee 2006. Two new species of skinks (Genus Sphenomorphus Fitzinger 1843) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History 9 (2): 151-162. Grismer, L. Lee. 2008. A new species of insular skink (Genus Sphenomorphus Fitzinger 1843) from the Langkawi Archipelago, Kedah, West Malaysia with the first report of the herpetofauna of Pulau Singa Besar and an updated checklist of the herpetofauna of Pulau Langkawi. Zootaxa 1691: 53-66 - get paper here GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species. Zootaxa 4092 (2): 231–242 - get paper here Grismer, L.L. 2007. A new species of small montane forest floor skink (genus Sphenomorphus Fitzinger 1843) form Southern peninsular Malaysia. Herpetologica 63 (4): 544-551 - get paper here Grismer, L.L.; Ahmad, N. & Kin Onn, C. 2009. A new, diminutive, upland Sphenomorphus Fitzinger 1843 (Squamata: Scincidae) from the Belum-Temengor Forest Complex, Peninsular Malaysia. Zootaxa 2312: 27–38 - get paper here Grismer, L.L.; Wood Jr., P.L. & Grismer, J.L. 2009. A New Insular Species of Skink of the Genus Sphenomorphus Strauch 1887 (Squamata: Scincidae) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. Tropical Life Sciences Research, 20(1): 51–69 - get paper here Inger, R.F.; Lian, R.F.; Lakim, M. & Yambun, P. 2001. New species of the lizard genus Sphenomorphus (Lacertilia: Scincidae), with notes on ecological and geographic distribution of species in Sabah, Malaysia. Raffles Bull. Zool. 49 (2): 181-189 - get paper here Lidth DE JEUDE, T.W. VAN 1905. Zoological results of the Dutch Scientific Expedition to Central-Borneo. The reptiles. Notes from the Leyden Museum 25 (4): 187-202. - get paper here Linkem, C.W.; Diesmos, A.C.; Brown, R.M. 2010. A NEW SPECIES OF SCINCID LIZARD (GENUS SPHENOMORPHUS) FROM PALAWAN ISLAND, PHILIPPINES. Herpetologica 66 (1): 67–79 - get paper here Linkem, Charles W.; Arvin C. Diesmos, Rafe M. Brown 2011. Molecular systematics of the Philippine forest skinks (Squamata: Scincidae: Sphenomorphus): testing morphological hypotheses of interspecific relationships. Zoological Journal of the Linnean Society 163: 1217–1243 - get paper here Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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