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Tytthoscincus jaripendek GRISMER, WOOD, QUAH, ANUAR, NGADI, MOHD-IZAM & AHMAD, 2017

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Common NamesE: Cameron highlands Forest skink
Malay: Mengkarung hutan Cameron highlands 
SynonymTytthoscincus jaripendek GRISMER, WOOD, QUAH, ANUAR, NGADI, MOHD-IZAM & AHMAD 2017 
DistributionPeninsular Malaysia (Pahang)

Type locality: Robinson’s Falls, Cameron Highlands, Pahang, Peninsular Malaysia (N 04°27.959′, E 101°23.129′; 1411 m in elevation  
TypesHolotype: LSUHC 11680, Adult male, collected by Evan S. H. Quah and L. Lee Grismer on 26 April 2014. Paratypes: LSUHC 11679, 11681 and 11987 all bear the same collection data as the holotype except the lat- ter was collected on 4 September 2014. 
DiagnosisDiagnosis: Tytthoscincus jaripendek sp. nov. can be differentiated from all other species of Tytthoscincus in the montane clade by having the combination of 8`1 superciliaries; a shallow, pigmented tympanic depression; no enlarged pectoral scales; 28–30 mid- body scales (MB); 63–65 paravertebral scales; 60–62 ventral scales; keeled, subdigital lamellae; five or six subdigital lamellae on the third finger; nine or ten subdigital lamellae on the fourth toe; TD/HL = 0.11– 0.13; HDL/SVL = 0.18–0.19; AXG/SVL = 0.55–0.57; FL/SVL = 0.18–0.21; HDL/SVL = 0.31–0.33 and a maximum SVL of 33.9 mm (Table 6). All non-ratiomet- ric characters are presented in all other Tytthoscincus and species of Sphenomorphus suspected of being Tytthoscincus in Grismer et al. (2016a: 237) and Karin et al. (2016: 416).

Comparisons: Tytthoscincus jaripendek sp. nov. can be differentiated from all other species of Tytthoscincus of the montane clade (Table 6), except its sister species T. kakikecil sp. nov., by having a shallow, pigmented tympanic depression vs. a well-developed, deeply recessed, non-pigmented tympanum. From T. kakikecil sp. nov., it differs discretely and statistically by having 63–65 vs. 67–73 paravertebral scales, 60–62 vs. 65–69 ventral scales (Tables 6, 7), and statistically significant differences in the mean number of paravertebral scales, ventral scales, 3TL and FL/SVL (Tables 6, 7). From all other species except T. perhentianensis, it differs further in having fewer midbody scales (28–30 vs. 30–38 col- lectively). Tytthoscincus jaripendek sp. nov. also bears statistically significant mean differences of all ten char- acters in varying combinations across all other species (Table 7). Meristic and body shape trends of these char- acters among species in the montane clade are presented in Figures 4 and 5. 
CommentHabitat: leaf-litter in hill dipterocarp forests. All specimens were collected by raking leaves that had accumulated at the foot of an earthen bank. Specimens were collected both day and night and during periods of heavy rain. 
EtymologyThe specific epithet jaripendek is derived from the Malay words jari meaning finger and pendek meaning short and refers to this species’ short digits as indicated by its low numbers of subdigital lamellae. 
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., EVAN S.H. QUAH, SHAHRUL ANUAR, EHWAN B. NGADI, NUR AMALINA MOHD IZAM and NORHAYATI AHMAD 2017. Systematics, ecomorphology, cryptic speciation and biogeography of the lizard genus Tytthoscincus Linkem, Diesmos & Brown (Squamata: Scincidae) from the sky- island archipelago of Peninsular Malaysia. Zoological Journal of the Linnean Society XX: 1–37 - get paper here
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