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Tytthoscincus keciktuek GRISMER, WOOD JR, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL & QUAH, 2018

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Higher TaxaScincidae, Sphenomorphinae, Scincoidea, Sauria, Squamata (lizards) 
Subspecies 
Common NamesE: Sungai Peres Forest Skink
Malay: Mengkarung Hutan Sungai Peres 
SynonymTytthoscincus keciktuek GRISMER, WOOD JR, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL4& QUAH 2018 
DistributionPeninsular Malaysia (Terengganu)

Type locality: small tributary of the Sungai Peres, Sekayu, Hulu Terengganu, Terengganu State, Peninsular Malaysia (4.9596° N, 102.9596° E; 74 m in elevation  
Reproduction 
TypesHolotype: LSUHC 13859, Adult female, collected by Murni- Azima, Amirrudin B. Ahmad, and M. Afiq-Shuhaimi. on 17 September 2017. 
DiagnosisDiagnosis. Tytthoscincus keciktuek sp. nov. can be differentiated from all other species of Tytthoscincus in the upland clade by having the combination of 7`1, superciliaries; a deeply set, unpigmented tympanum; enlarged pectoral scales; 30 midbody scale rows; 66 paravertebral scales; 60 ventral scales; keeled, subdigital lamellae; seven subdigital lamellae on the third finger; 11 subdigital lamellae on the fourth toe; EO/HL = 0.12; HL/SVL = 0.21; AXG/SVL = 0.54; FL/SVL = 0.25; HDL/SVL = 0.37; and a maximum SVL of 36.2 mm (Tables 5,6). All non-ratiometric characters are scored across all other Tytthoscincus and species of Sphenomorphus suspected of being Tytthoscincus in Grismer et al. (2016: 237) and Karin et al. (2016: 416).

Comparisons. Tytthoscincus keciktuek sp. nov. is most closely related to the undescribed species from the Tembak Reservoir area from which it differs by an uncorrected pairwise sequence divergence of 5.3%. Unfortunately, the Tembat specimen was so badly damaged by insects in a pitfall trap that no morphological data were unobtainable. Differences between T. keciktuek sp. nov. and T. monticolus sp. nov. are presented above. Differences between T. keciktuek sp. nov. and all other species of the upland clade are presented in Table 6 in Grismer et al. 2018. 
CommentHabitat: leaf-litter along riparian systems in lowland dipterocarp forest 
EtymologyThe name "keciktuek", is a colloquial word in the local dialect of rural Terengganu that means “small, small things” and refers to all small animmals. It is used here in reference to the small size of all the species of the upland clade of Tytthoscincus. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species Zootaxa 4092 (2): 231–242 - get paper here
  • GRISMER, L. LEE; PERRY L. JR. WOOD, AMIRRUDIN B. AHMAD, B. S. BAIZUL-HAFSYAM, M. AFIQ-SHUHAIMI, SYED A. RIZAL, EVAN S. H. QUAH 2018. Two new Tytthoscincus Linkem, Diesmos, & Brown (Squamata; Scincidae) from Peninsular Malaysia and another case of microsyntopy between ecologically specialized, unrelated, leaf-litter species. Zootaxa 4425 (1): 087–107 - get paper here
  • KARIN, BENJAMIN R.; INDRANEIL DAS, AARON M. BAUER 2016. Two new species of diminutive leaf-litter skinks (Squamata: Scincidae: Tytthoscincus) from Gunung Penrissen, Sarawak, Malaysia (northern Borneo). Zootaxa 4093 (3): 407–423
 
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