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Tytthoscincus monticolus GRISMER, WOOD, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL & QUAH, 2018

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Higher TaxaScincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Sungai Bubu Forest Skink
Malay: Mengkarung Hutan Sungai Bubu 
SynonymTytthoscincus monticolus GRISMER, WOOD, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL4& QUAH 2018 
DistributionPeninsular Malaysia (Terengganu)

Type locality: hilly area near Sungai Bubu, Sekayu, Hulu Terengganu, Terengganu State, Peninsular Malaysia (4.9710° N, 102.9531° E; 174 m in elevation  
Reproduction 
TypesHolotype: LSUHC 13858, Adult male, collected by M. Afiq-Shuhaimi and Baizul-Hafsyam B. S. on 5 August 2017. 
DiagnosisDiagnosis Tytthoscincus monticolus sp. nov. can be differentiated from all other species of Tytthoscincus in the upland clade by having the combination of 8`1, superciliaries; a deeply set, unpigmented tympanum; enlarged pectoral scales; 31 midbody scale rows; 69 paravertebral scales; 68 ventral scales; keeled, subdigital lamellae; seven subdigital lamellae on the third finger; 12 subdigital lamellae on the fourth toe; EO/HL = 0.16; HL/SVL = 0.17; AXG/SVL = 0.55; FL/SVL = 0.22; HDL/SVL = 0.33; and a maximum SVL of 36.6 mm (Tables 5,6). All characters are scored across all other Tytthoscincus and species of Sphenomorphus suspected of being Tytthoscincus in Grismer et al. (2016a:237) and Karin et al. (2016:416).


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CommentHabitat: leaf-litter in hilly areas along riparian systems in lowland dipterocarp forest 
EtymologyThe specific epithet monticolus, comes from the Latin monti meaning mountain and colus meaning dweller of or within and refers to this species inhabiting hilly areas. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species. Zootaxa 4092 (2): 231–242 - get paper here
  • GRISMER, L. LEE; PERRY L. JR. WOOD, AMIRRUDIN B. AHMAD, B. S. BAIZUL-HAFSYAM, M. AFIQ-SHUHAIMI, SYED A. RIZAL, EVAN S. H. QUAH 2018. Two new Tytthoscincus Linkem, Diesmos, & Brown (Squamata; Scincidae) from Peninsular Malaysia and another case of microsyntopy between ecologically specialized, unrelated, leaf-litter species. Zootaxa 4425 (1): 087–107 - get paper here
  • KARIN, BENJAMIN R.; INDRANEIL DAS, AARON M. BAUER 2016. Two new species of diminutive leaf-litter skinks (Squamata: Scincidae: Tytthoscincus) from Gunung Penrissen, Sarawak, Malaysia (northern Borneo). Zootaxa 4093 (3): 407–423 - get paper here
  • Syafiq, Muhamad Fatihah; Baizul Hafsyam Badli-Sham, Noor-Shahirah Ibrahim, Lukman H. B. Ismail, Muhammad Afif Mohd Amin, Goh Shu Xian, Rabiatul ‘Aqilah Mohamad Ariffin, Muhammad Afiq-Suhaimi, Lok Kar Men, Dila Adilina Danelo, Mohamad Aqmal-Naser, Muh 2024. TAXONOMIC COMPOSITION, DIVERSITY, AND CONSERVATION STATUS OF REPTILIAN FAUNA AT SEKAYU LOWLAND FORESTS, TERENGGANU, PENINSULAR MALAYSIA. Russian Journal of Herpetology Vol. 31, No. 1, 2024, pp. 14 – 23 - get paper here
 
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