Tytthoscincus monticolus GRISMER, WOOD, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL & QUAH, 2018

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Higher Taxa	Scincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards)
Subspecies
Common Names	E: Sungai Bubu Forest Skink Malay: Mengkarung Hutan Sungai Bubu
Synonym	Tytthoscincus monticolus GRISMER, WOOD, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL4& QUAH 2018
Distribution	Peninsular Malaysia (Terengganu) Type locality: hilly area near Sungai Bubu, Sekayu, Hulu Terengganu, Terengganu State, Peninsular Malaysia (4.9710° N, 102.9531° E; 174 m in elevation
Reproduction
Types	Holotype: LSUHC 13858, Adult male, collected by M. Afiq-Shuhaimi and Baizul-Hafsyam B. S. on 5 August 2017.
Diagnosis	Diagnosis Tytthoscincus monticolus sp. nov. can be differentiated from all other species of Tytthoscincus in the upland clade by having the combination of 8`1, superciliaries; a deeply set, unpigmented tympanum; enlarged pectoral scales; 31 midbody scale rows; 69 paravertebral scales; 68 ventral scales; keeled, subdigital lamellae; seven subdigital lamellae on the third finger; 12 subdigital lamellae on the fourth toe; EO/HL = 0.16; HL/SVL = 0.17; AXG/SVL = 0.55; FL/SVL = 0.22; HDL/SVL = 0.33; and a maximum SVL of 36.6 mm (Tables 5,6). All characters are scored across all other Tytthoscincus and species of Sphenomorphus suspected of being Tytthoscincus in Grismer et al. (2016a:237) and Karin et al. (2016:416).
Comment	Habitat: leaf-litter in hilly areas along riparian systems in lowland dipterocarp forest
Etymology	The specific epithet monticolus, comes from the Latin monti meaning mountain and colus meaning dweller of or within and refers to this species inhabiting hilly areas.
References	GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here GRISMER, L. LEE; MOHD ABDUL MUIN, PERRY L. WOOD, JR., SHAHRUL ANUAR, & CHARLES W. LINKEM 2016. The transfer of two clades of Malaysian Sphenomorphus Fitzinger (Squamata: Scincidae) into the genus Tytthoscincus Linkem, Diesmos, & Brown and the description of a new Malaysian swamp-dwelling species. Zootaxa 4092 (2): 231–242 - get paper here GRISMER, L. LEE; PERRY L. JR. WOOD, AMIRRUDIN B. AHMAD, B. S. BAIZUL-HAFSYAM, M. AFIQ-SHUHAIMI, SYED A. RIZAL, EVAN S. H. QUAH 2018. Two new Tytthoscincus Linkem, Diesmos, & Brown (Squamata; Scincidae) from Peninsular Malaysia and another case of microsyntopy between ecologically specialized, unrelated, leaf-litter species. Zootaxa 4425 (1): 087–107 - get paper here KARIN, BENJAMIN R.; INDRANEIL DAS, AARON M. BAUER 2016. Two new species of diminutive leaf-litter skinks (Squamata: Scincidae: Tytthoscincus) from Gunung Penrissen, Sarawak, Malaysia (northern Borneo). Zootaxa 4093 (3): 407–423 - get paper here
External links	National Center for Biotechnology Information Google images

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