Tytthoscincus monticolus GRISMER, WOOD, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL & QUAH, 2018
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Tytthoscincus monticolus
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| Higher Taxa | Scincidae, Sphenomorphinae (Sphenomorphini), Scincoidea, Sauria, Squamata (lizards) |
| Subspecies | |
| Common Names | E: Sungai Bubu Forest Skink Malay: Mengkarung Hutan Sungai Bubu |
| Synonym | Tytthoscincus monticolus GRISMER, WOOD, AHMAD, BAIZUL-HAFSYAM, AFIQ-SHUHAIMI, RIZAL4& QUAH 2018 |
| Distribution | Peninsular Malaysia (Terengganu) Type locality: hilly area near Sungai Bubu, Sekayu, Hulu Terengganu, Terengganu State, Peninsular Malaysia (4.9710° N, 102.9531° E; 174 m in elevation |
| Reproduction | |
| Types | Holotype: LSUHC 13858, Adult male, collected by M. Afiq-Shuhaimi and Baizul-Hafsyam B. S. on 5 August 2017. |
| Diagnosis | Diagnosis Tytthoscincus monticolus sp. nov. can be differentiated from all other species of Tytthoscincus in the upland clade by having the combination of 8`1, superciliaries; a deeply set, unpigmented tympanum; enlarged pectoral scales; 31 midbody scale rows; 69 paravertebral scales; 68 ventral scales; keeled, subdigital lamellae; seven subdigital lamellae on the third finger; 12 subdigital lamellae on the fourth toe; EO/HL = 0.16; HL/SVL = 0.17; AXG/SVL = 0.55; FL/SVL = 0.22; HDL/SVL = 0.33; and a maximum SVL of 36.6 mm (Tables 5,6). All characters are scored across all other Tytthoscincus and species of Sphenomorphus suspected of being Tytthoscincus in Grismer et al. (2016a:237) and Karin et al. (2016:416). Comparisons. Tytthoscincus monticolus sp. nov. is most closely related to T. perhentianensis (Fig. 4) and differs from it by an uncorrected pairwise sequence divergence of 3.5%. It is differentiated from T. perhentianensis morphologically by having one vs. two loreal scales; 69 vs. 65 or 66 paravertebral scales; 68 vs. 61 or 62 ventral scales; seven vs. six subdigital lamellae on the third finger; and 12 vs. 10 subdigital lamellae on the fourth toe. It differs further by having a larger ear opening (0.16 vs 0.13–0.14 EO/HL); a shorter head (0.17 vs. 0.22–0.23 HL/ SVL); longer hind limbs (0.33 vs. 0.29–0.30 HDL/SVL); and a larger maximum SVL (36.2 mm vs. 30.0 mm: Table 6). It differs from T. keciktuek (see description below) with which it is likely syntopic by an uncorrected pairwise sequence divergence of 9.8% and having one vs. two loreals; 68 vs 60 ventral scale rows; a larger ear opening (0.16 vs 0.12 EO/HL); a shorter head (0.17 vs. 0.21 HL/SVL); shorter forelimbs (0.22 vs. 0.25 FL/SVL); shorter hind limbs (0.33 vs. 0.37 HDL/SVL); and a dorsal pattern bearing white speckling vs. being nearly unicolor (Table 5; Fig. 6). Differences between T. monticolus sp. nov. and all other species of the upland clade are presented in Table 6 in Grismer et al. 2018. |
| Comment | Habitat: leaf-litter in hilly areas along riparian systems in lowland dipterocarp forest |
| Etymology | The specific epithet monticolus, comes from the Latin monti meaning mountain and colus meaning dweller of or within and refers to this species inhabiting hilly areas. |
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