Uma inornata COPE, 1895

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Higher Taxa	Phrynosomatidae, Phrynosomatinae, Callisaurini; Iguania, Sauria, Squamata (lizards)
Subspecies
Common Names	E: Coachella Valley Fringe-toed Lizard
Synonym	Uma notata inornata COPE 1895: 939 Uma notata inornata — NORRIS 1958: 289 Uma inornata — MAYHEW 1964 Uma inornata — STEBBINS 1985: 119 Uma inornata — CROTHER 2000: 54 Uma inornata — COLLINS & TAGGART 2009 Uma inornata — GOTTSCHO et al. 2016 Uma inornata — HANSEN & SHEDD 2025
Distribution	USA (SE California: Coachella Valley, Riverside County) Type locality: "Colorado Desert, San Diego County, California" (in errore, doubtless from Riverside County, California fide HEIFETZ 1941).
Reproduction	oviparous
Types	Holotype: lost, was USNM 16500 (lost fide HEIFETZ 1941), collected by C.R. Orcutt.
Diagnosis	Diagnosis: Abdomen immaculate. Similar to U. n. notata in all other characters. Gular markings like those of U. n. notata; usually three internasal rows; femoral pores 18 to 28; fringe scales on fourth toe 25 to 36; no reddish brown in dorsal pattern; dorsal ocelli arranged in semilineate pattern as in U. n. notata (from HEIFETZ 1941).
Comment	Cope (1895) described U. inornata from a single specimen. Not listed by Collins 1997. Trépanier & Murphy (2001) found that the preferred phylogeny had two primary clades: one consisting of U. scoparia and the other placing U. inornata inside the clade containing U. notata. Uma inornata was most closely related to nearby U. notata notata, as opposed to more distant U. notata rufopunctata. Distribution: see map in Gottscho et al. 2016: (Fig. 1), Derycke et al. 2020: 68 (Fig. 1).
Etymology	Named after Latin inornata, unadorned. [“...No black spots on belly or crescents on throat...”].
References	Collins, J.T. and T. W. Taggart 2009. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Sixth Edition. Center for North American Herpetology, 48 pp. Cope, E.D. 1895. On the species of Uma and Xantusia. American Naturalist 29: 938-939 - get paper here Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92 DERYCKE, ELIZABETH G.; ANDREW D. GOTTSCHO, ANDREW D. GOTTSCHO, DANIEL G. MULCAHY, KEVIN DE QUEIROZ 2020. A new cryptic species of fringe-toed lizards from southwestern Arizona with a revised taxonomy of the Uma notata species complex (Squamata: Phrynosomatidae). Zootaxa 4778 (1): 67–100 - get paper here Durtsche, R.D. 1995. Foraging ecology of the fringe-toed lizard, Uma inornata, during periods of high and low food abundance Copeia 1995 (4): 915-926. - get paper here Fisher, M., A. Muth & R.E. Johnson 2020. A Long-term Study of Home Range of Coachella Fringe-Toed Lizards, Uma inornata Journal of Herpetology 54 (2): 174-182. - get paper here Gottscho, A.D., Wood, D.A., Vandergast, A.G., Lemos-Espinal, J., Gatesy, J., Reeder, T.W. 2016. Lineage Diversification of Fringe-toed Lizards (Phrynosomatidae: Uma notata complex) in the Colorado Desert: Delimiting Species in the Presence of Gene Flow. Molecular Phylogenetics and Evolution 106: 103-117 - get paper here Hansen, Robert and Jackson D. Shedd 2025. California Amphibians and Reptiles. Princeton University Press, 528 pp. - get paper here Hedtke, Shannon M., Kelly R. Zamudio, Christopher A. Phillips, Jonathan Losos and Phil Brylski 2007. Conservation genetics of the endangered Coachella Valley fringe-toed lizard, Uma inornata. Herpetologica 63 (4): 411-420 - get paper here Jones, L.L. & Lovich, R.E. 2009. Lizards of the American Southwest. A photographic field guide. Rio Nuevo Publishers, Tucson, AZ, 568 pp. [review in Reptilia 86: 84] - get paper here Mayhew, W.W. 1964. Photoperiodic responses in three species of the lizard genus Uma. Herpetologica 20 (2): 95-113. - get paper here Mayhew, W.W. 1964. Taxonomic Status of California Populations of the Lizard Genus Uma. Herpetologica 20 (3): 170-183. - get paper here Mayhew, W.W. 1965. Reproduction in the Sand-Dwelling Lizard Uma inornata. Herpetologica 21 (1): 39-55. - get paper here MUTH, ALLAN, MARK FISHER & CHRISTOPHER R. TRACY. 2023. A Long-Term Study of Coachella Fringe-Toed Lizards, Uma inornata, Reveals Seasonal and Annual Variation in Size and Growth. Journal of Herpetology 57(2): 229–237. - get paper here Pilliod, D., Jeffries, M., Arkle, R. and Olson, D. 2024. Climate Futures for Lizards and Snakes in Western North America May Result in New Species Management Issues. Ecol Evol, 14: e70379 - get paper here Pough, F. Harvey 1973. Uma inornata. Catalogue of American Amphibians and Reptiles (126): 1-2 - get paper here Smith, Hobart M. 1946. Handbook of Lizards: Lizards of the United States and of Canada. Comstock, Ithaca, NY, xxii + 557 pp. Stebbins,R.C. 1985. A Field Guide to Western Reptiles and Amphibians, 2nd ed. Houghton Mifflin, Boston Trépanier, T.L.; Murphy, R.W. 2001. The Coachella Valley fringe-toed lizard (Uma inornata): genetic diversity and phylogenetic relationships of an endangered species. Molecular Phylogenetics and Evolution 18(3):327-334 - get paper here Turner, F.B., D.C. Weaver & J.C. Rorabaugh 1984. Effects of Reduction in Windblown Sand on the Abundance of the Fringe-Toed Lizard (Uma inornata) in the Coachella Valley, California Copeia 1984 (2): 370-378. - get paper here Werning, Heiko 2012. Die Reptilien und Amphibien des Südwestens. Draco 13 (50): 18-60 - get paper here
External links	IUCN Red List National Center for Biotechnology Information http://blackbook.org/~sullivan/indexes/sciIndexS-Z.html http://www.californiaherps.com/lizards/pages/u.inornata.html Google images

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