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Varanus gouldii GRAY, 1838

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Higher TaxaVaranidae, Platynota, Anguimorpha, Sauria, Squamata (lizards) 
SubspeciesVaranus gouldii gouldii (GRAY 1838)
Varanus gouldii flavirufus MERTENS 1958 
Common NamesE: Sand Monitor, Gould's Goanna
G: Gouldswaran 
SynonymVaranus gouldii gouldii (GRAY 1838)
Hydrosaurus gouldii GRAY 1838: 394
Monitor gouldii — SCHLEGEL 1839: 78
Varanus gouldii — DUMÉRIL & DUMÉRIL 1851: 52
Varanus gouldii — BOULENGER 1885: 320
Varanus gouldi — DE ROOIJ 1915: 149
Varanus (Varanus) gouldii - MERTENS 1942
Varanus gouldii gouldii — MERTENS 1958
Varanus panoptes STORR 1980: 273
Pantherosaurus barryjonesi WELLS & WELLINGTON 1985: 21
Varanus gouldii gouldii — BÖHME 1991: 38
Varanus gouldii gouldii — DE LISLE 1996: 126
Varanus gouldii gouldii — COGGER 2000: 368
Varanus gouldii — WILSON & SWAN 2010
Varanus (Varanus) gouldii — BUCKLITSCH et al. 2016: 50

Varanus gouldii flavirufus MERTENS 1958
Hydrosaurus gouldii GRAY 1838: 394
Varanus gouldii DUMÉRIL 1851: 52
Varanus gouldii flavirufus MERTENS 1958: 229
Varanus flavirufus flavirufus — BÖHME 1991: 38
Varanus gouldii flavirufus — COGGER 2000: 368
Varanus gouldii flavirufus — BÖHME 2002
Varanus flavirufus — DE LISLE 2009: 17 
DistributionAustralia (except arid interior, including Queensland, New South Wales), S New Guinea

flavirufus: Central Australia in the area of the red soils; westwards to the Canning Stock Route (Western Australia); Type locality: Bat Caves near Alice Springs, Northern Territory.

Type locality: "Australia", restricted by Opinion 1948 of the International Commission of Zool. Nomenclature (2000), due to the designation of a neotype, to Karakatta, Perth, Western Australia. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesNeotype: BMNH 1997.1 (designated by the Comission of Zoological Nomenclature in March 2000, Opinion 1948), from Karrakatta, Perth, Western Australia; see comment.
Holotype: SMF 53271 [flavirufus] 
CommentSubspecies: Varanus gouldii rubidus is listed here as a subspecies of V. panoptes following COGGER 2000. Varanus gouldii horni BÖHME 1988 is not listed by COGGER 2000.

Types: The lectotype of V. gouldii has been a specimen of V. panoptes, formally rendering the name panoptes a junior synonym of gouldii. Sprackland et al. (1997) proposed that the lectotype of gouldii be set aside and a neotype designated in accord with accustomed usage. The International Commission of Zoological Nomenclature (2000: Opinion 1948) accepted an application to set aside the lectotype and to replace it by a neotype fitting with what was understood to be V. gouldii before.

Illustration in Schmida (2000) and Bennet (2003).

Synonymy: partly after SHEA & SADLIER 1999. 
References
  • Bennet, D.F. 2003. Australian Monitors. Reptilia (GB) (30): 12-19 - get paper here
  • Bennet, D.F. 2003. Der Varanus-gouldii-Komplex. Reptilia (Münster) 8 (43): 26-28 - get paper here
  • Bennet, D.F. 2003. Australische Warane. Reptilia (Münster) 8 (43): 18-25 - get paper here
  • Bennet, D.F. 2003. The Varanus gouldii group. Reptilia (GB) (30): 27-29 - get paper here
  • Böhme, W. 1988. Der Arguswaran (Varanus panoptes, Storr 1980) auf Neuguinea: Varanus panoptes horni spp. n. Salamandra 24 (2-3): 87-101 - get paper here
  • Böhme, W. 1991. The identity of Varanus gouldii (Gray, 1838), and the nomenclature of the V. gouldii species complex. Mertensiella, 2, 38-41.
  • Böhme,W. 2003. Checklist of the living monitor lizards of the world (family Varanidae). Zoologische Verhandelingen 341: 6-43 - get paper here
  • Böhme,W. & T. ZIEGLER 1998. Comments on the proposed conservation of the names Hydrosaurus gouldii Gray, 1838 and Varanus panoptes STORR, 1980 (Reptilia, Squamata) by the designation of a neotype for Hydrosaurus gouldii (Case 3042; see BZN 54: 95-99, 249-250; 55: 106-111). Bull. Zool. Nomenclature 55 (3): 173-174. - get paper here
  • Bonetti, Mathilde 2002. 100 Sauri. Mondadori (Milano), 192 pp.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp.
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • De Lisle, H.F. 1996. Natural History of Monitor Lizards. Krieger, Malabar (Florida)
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp.
  • Duméril, A.M.C. & A. H. A. Duméril 1851. Catalogue méthodique de la collection des reptiles du Muséum d'Histoire Naturelle de Paris. Gide et Baudry/Roret, Paris, 224 pp.
  • Earley, Ryan L.; Omar Attum, Perri Eason 2002. Varanid combat: perspectives from game theory. Amphibia-Reptilia 23 (4): 469-485 - get paper here
  • FRÝDLOVÁ, PETRA and DANIEL FRYNTA 2010. A test of Rensch’s rule in varanid lizards. Biological Journal of the Linnean Society 100: 293–306 - get paper here
  • Gray, J. E. 1838. Catalogue of the slender-tongued saurians, with descriptions of many new genera and species. Part 3. Ann. Mag. Nat. Hist. (1) 1: 388-394 - get paper here
  • Grundke, Frank D.;Grundke, Birgit 1992. Feldherpetologische Beiträge zur Fauna Australiens. Teil 3: Zur Kenntnis des Varanus gouldii-Komplexes in Northern Territory und Queensland. Herpetofauna 14 (81): 11-19 - get paper here
  • ICZN 2000. OPINION 1948. Hydrosaurus gouldii Gr ay, 1838 (currently Varanus gouldii) and Varanus panoptes Storr, 1980 (Reptilia, Squamata): specific names conserved by the designation of a neotype for H. gouldii. Bull. Zool. Nomenclature 57 (1) - get paper here
  • Limpus, Duncan J. 1995. Observations of Varanus gouldii (Varanidae) at Mon Repos Beach, Bundaberg. Herpetofauna (Sydney) 25 (2): 14-16
  • McCurry MR, Mahony M, Clausen PD, Quayle MR, Walmsley CW, Jessop TS, et al. 2015. The Relationship between Cranial Structure, Biomechanical Performance and Ecological Diversity in Varanoid Lizards. PLoS ONE 10(6): e0130625. doi:10.1371/journal.pone.0130625 - get paper here
  • Mertens, R. 1942. Die Familie der Warane (Varanidae), 3. Teil: Taxonomie. Abh. Senckenb. naturf. Ges., 466: 235-391
  • Mertens, R. 1958. Bemerkungen über die Warane Australiens. Senckenbergiana biologica 39: 229-264
  • Michael, D.R.; D.B. Lindenmayer; M. Crane; C. MacGregor; R. Montague-Drake; L. McBurney. 2011. Reptilia, Murray catchment, New South Wales, southeastern Australia. Check List 7 (1):25-29 - get paper here
  • Pianka, E. R. 1969. Habitat specificity, speciation, and species density in Australian desert lizards. Ecology 50 (3): 498-502 - get paper here
  • Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press, 599 pp - get paper here
  • Pianka, E.R. 2003. Australian Desert Varanids. Reptilia (GB) (30): 20-26 - get paper here
  • Pianka, E.R. 2003. Die Warane der australischen Wüste. Reptilia (Münster) 8 (43): 29-35 - get paper here
  • Read, John L. and Vince Scoleri. 2015. Ecological Implications of Reptile Mesopredator Release in Arid South Australia. Journal of Herpetology 49 (1): 64-69
  • Rese, R. 1988. Beitrag zur Lebenserwartung von Grosswaranen in Gefangenschaft. Sauria 10 (1): 25-26 - get paper here
  • Sarre, Kazumi Matsubara; Stephen D.; Arthur Georges, Yoichi Matsuda, Jennifer A. Marshall Graves; Tariq Ezaz 2014. Highly Differentiated ZW Sex Microchromosomes in the Australian Varanus Species Evolved through Rapid Amplification of Repetitive Sequences. Plos One, DOI: 10.1371/journal.pone.0095226 - get paper here
  • Schlegel, H. 1839. Abbildungen neuer oder unvollständig bekannter Amphibien, nach der Natur oder dem Leben entworfen und mit einem erläuternden Texte begleitet. Arne and Co., Düsseldorf, xiv + 141 pp.
  • Schmida, G. 2000. Exkursionen bei Mackay und Proserpine. DATZ 53 (9): 8-12
  • Shea, G. M. 1994. Three species of goanna occur in the Sydney Basin. Herpetofauna (Sydney) 24 (2): 14-18
  • Sprackland R G.; Smith H M. & Strimple P D. 1997. Hydrosaurus gouldii Gray, 1838 (currently Varanus gouldii) and Varanus panoptes Storr, 1980 (Reptilia, Squamata): Proposed conservation of the specific names by the designation of a neotype for H. gouldii. Bull. Zool. Nomenclature 54 (2): 95-99 - get paper here
  • Sternfeld, R. 1925. Beiträge zur Herpetologie Inner-Australiens. Abhandlungen Herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft, 38: 221—251
  • Storr G M 1980. The monitor lizards (genus Varanus Merrem, 1820) of Western Australia. Rec. West. Aust. Mus. 8 (2): 237-293 - get paper here
  • Sutherland, Duncan R 2011. Dietary Niche Overlap and Size Partitioning in Sympatric Varanid Lizards. Herpetologica 67 (2): 146-153. - get paper here
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
  • Ziegler, Thomas; Böhme, Wolfgang 1996. Neue Erkenntnisse zur Geschlechtsunterscheidung bei squamaten Reptilien. Kleintierpraxis 41 (8): 585-590
 
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