Vipera lotievi NILSON, TUNIYEV, ORLOV, HOGGREN & ANDREN, 1995

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Higher Taxa	Viperidae, Viperinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names	E: Lotiev’s viper G: Lotievs Otter Russian: Гадюка Лотиева
Synonym	Vipera lotievi NILSON et al. 1995 Vipera berus — (fide KHALIKOV, pers. comm.) Vipera ursini renardi — (fide KHALIKOV, pers. comm.) Vipera (Pelias) lotievi — NILSON et al. 1999: 103 Vipera lotievi — MCDIARMID, CAMPBELL & TOURÉ 1999: 406 Vipera renardi lotievi — DELY & JOGER 2005 Pelias lotievi — WALLACH et al. 2014: 544 Pelias lotievi — BEKOSHVILI & DORONIN 2015 Pelias lotievi — TUNIYEV 2016 Pelias lotievi — ISKENDEROV et al. 2017 Pelias lotievi — DUNAEV & ORLOVA 2017 Pelias lotievi — TUNIYEV et al. 2019: 147
Distribution	Russia (northern slope of Caucasus), Azerbaijan Type locality: S Russia (N Caucasus), Republic of Georgia
Reproduction	ovovivparous
Types	Holotype: ZISP (ZIL) 20309.
Diagnosis
Comment	Venomous! Synonymy: V. lotievi is nested within V. renardi (F. Martínez-Freiría, pers. comm., 5 Sep 2019, SEH)
Etymology	Named after K. Yu Lotiev, a Russian herpetologist who collected the viper holotype (1986).
References	Bekoshvili, David and Igor V Doronin. 2015. New data on the distribution of snakes in Georgia (Causasus). Herpetological Review 46 (3): 388-390 - get paper here Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Dunaev E.A., Orlova V.F. 2017. Amphibians and reptiles of Russia. Atlas and determination. 2nd. ed. Moscow: Phyton XXI, 328 p Iskenderov, T. M.; S. B. Akhmedov, and S. N. Bunyatova 2017. LOTIEV’S VIPER (PELIAS LOTIEVI, SERPENTES, VIPERIDAE), A SPECIES NEW TO THE FAUNA OF AZERBAIJAN REPUBLIC. Zoologichesky zhurnal 96 (1): 121–124 Mallow, D. Ludwig, D. & Nilson, G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Krieger, Malabar, Florida, 410 pp. [review in HR 35: 200, Reptilia 35: 74] McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp. Nilson G; Tuniyev B S; Orlov N; Hoggren M; Andren C 1995. Systematics of the vipers of the Caucasus: Polymorphism or sibling species? Asiatic Herpetological Research 6: 1-16 - get paper here Nilson,G.; TUNIYEV,B.; ANDRÉN,C. & ORLOV,N. 1999. Vipers of Caucasus: Taxonomic considerations. Kaupia (Darmstadt) (8): 103-106 Phelps, T. 2010. Old World Vipers. Edition Chimaira, Frankfurt, 558 pp. [critical review in Sauria 33 (3): 19 and HR 43: 503] Tuniyev B.S. 2016. Rare species of shield-head vipers in the Caucasus. Nature Conservation Research 1 (3): 11–25 - get paper here Tuniyev, B.; Nilson, G. & Andrén, C. 2010. A new species of viper (Reptilia, Viperidae) from the Altay and Saur Mountains, Kazakhstan. Russ. J. Herpetol. 17 (2): 110-120 - get paper here Tuniyev, B.S.; N.L. Orlov, N.B. Ananjeva and A.L. Aghasyan 2019. Snakes of the Caucasus: taxonomic diversity, distribution, conservation. St. Petersburg, Moscow. KMK Scientific Press. 2019. 276 pp. - get paper here Tuniyev, S. B.; N. Orlov, B. Tuniyev, A. Kidov 2013. On the Taxonomical Status of Steppe Viper from Foothills of the South Macroslope of the East Caucasus. Russ. J. Herpetol. 20 (2): 129-146 - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
External links	IUCN Red List National Center for Biotechnology Information http://www.kingsnake.com/viperidae/images/photos.htm http://www.wildherps.com/species/E.tigrinus.html Google images

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