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Vipera walser GHIELMI, MENEGON, MARSDEN, LADDAGA & URSENBACHER, 2016

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Higher TaxaViperidae, Viperinae, Colubroidea, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesG: Piemont-Viper 
SynonymVipera walser GHIELMI, MENEGON, MARSDEN, LADDAGA & URSENBACHER 2016 
DistributionNW Italy (Pennine Alps)

Type locality: San Giovanni d’Andorno, strada per Oropa at 1300 m elevation in the Alps north of town of Biella, a subrange of the Pennine Alps, north-western Italy.  
Reproductionviviparous 
TypesHolotype: MSNG 34485, Adult female, collected in S. Giovanni d’Andorno, on the road to Oropa in the Biella prealps, at about 1300 m a.s.l. by A. Rosazza in the summer of 1930 (Fig. 5).
Paratypes: One adult male: MSNG33638M collected at Monte Rosso del Croso, on 30 August 1933. One juvenile male: MSNG33637B and one subadult male: MSNG30818C collected at Alpe Finestre by Felice Capra, respectively, on 28 July 1930 and 15 August 1928. One adult female: MSNG30818A, one subadult female: MSNG30818B, and two juvenile females: MSNG33637C and MSNG33637D collected by Felice Capra at Alpe Finestre between August 1928 and August 1939. One juvenile female: MSNG30286 collected by F. Capra at Monte Rosso del Croso on 12 September 1934; one adult female MSNG33637A col- lected by F. Capra at Alpe le Piane on 5 August 1937; one adult female MSNG41663 collected by A. Margiocco at Piedicavallo in September 1967. 
CommentDistribution: see map in Ghielmi et al. 2016: 168 (Fig. 8)

Phylogenetics: V. walseri nests within dinniki, kaznakovi, darevskii, ursinii, eriwanensis. Not closely related to V. aspis or V. berus. See tree in Ghielmi et al. 2016: 163 (Fig. 1). 
References
  • Ghielmi, S., Menegon, M., Marsden, S. J., Laddaga, L. and Ursenbacher, S. 2016. A new vertebrate for Europe: the discovery of a range-restricted relict viper in the western Italian Alps. Journal of Zoological Systematics and Evolutionary Research, 54: 161–173.doi: 10.1111/jzs.12138 - get paper here
  • Trapp, Benny 2017. Auf der Suche nach einer neubeschriebenen Viper in den italienischen Alpen: die Piemont-Viper (Vipera walser) Terraria-Elaphe 2017 (1): 78-83 - get paper here
 
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