You are here » home advanced search Zamenis longissimus

Zamenis longissimus (LAURENTI, 1768)

IUCN Red List - Zamenis longissimus - Least Concern, LC

Can you confirm these amateur observations of Zamenis longissimus?

Add your own observation of
Zamenis longissimus »

Find more photos by Google images search: Google images

Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Aesculapean Snake
G: Äskulapnatter
Croatian: bjelica
Russian: Эскулапов полоз
Italian: saettone comune 
SynonymNatrix longissima LAURENTI 1768: 74
Coluber aesculapi LACEPEDE 1789
Elaphis Aesculapii — DUMÉRIL, BIBRON & DUMÉRIL 1854: 278
Coluber esculapii — BOETTGER 1889: 271
Coluber esculapii — SCLATER 1891: 33
Coluber longissimus — BOULENGER 1894: 52
Coluber Aesculapii — ZANG 1905
Coluber longissimus — BOULENGER 1913
Elaphe longissima — ENGELMANN et al 1993
Elaphe longissima — SCHULZ 1996
Elaphe longissima — STOEV 2000
Zamenis longissimus — UTIGER et al. 2002
Zamenis longissimus — SCHLÜTER 2006
Zamenis longissima — VENCHI & SINDACO 2006
Zamenis longissimus — KWET 2007
Zamenis longissimus — WALLACH et al. 2014: 808 
DistributionNW Spain, France, Italy, S Switzerland, Andorra,
Germany (Taunus, Neckar river, Passau, Salzach-river, Berchtesgaden), SE Poland,
S Austria (except Tirol and Vorarlberg),
Czechoslovakia, Hungary, Romania, Bulgaria, N Turkey, Greece (incl. Corfu = Corfou),
Croatia (including some Adriatic islands), Slovenia, Bosnia, Hercegovina, Montenegro, Macedonia, Serbia, S Slovakia,
Albania, S Czech Republic, Republic of Georgia, NW Iran,
S Russia: south to Kuban river, SW Ukraine, S Moldova, Azerbaijan

Introduced to the United Kingdom (Wales) [fide arc-trust.org]

Type locality: Austria (Laurenti, 1768); Austria, Vienna (Mertens & Müller, 1928)  
Reproductionoviparous 
TypesType: nondesignated, present location unknown (fide V. Wallach, pers. comm.) 
DiagnosisAdditional details, e.g. a detailed description or comparisons (2249 characters), are available for collaborators and contributors. Please contact us for details. 
CommentElaphe longissima romana has been elevated to species status by Lenk & Wüster 1999 and is now called E. lineata.

Type species: Coluber aesculapi LACEPEDE 1789 is the type species of the genus Zamenis WAGLER 1830 (fide WILLIAMS & WALLACH 1989: 161).

Synonymy: Kaiser et al. 2013 considered the generic name Richardwellsus Hoser 2012 invalid and rejected its use instead of Zamenis.

Habitat: partly arboreal (Harrington et al. 2018).

Distribution: Not in Poland fide Geniez 2018, Salvi et al. 2018, but recorded from isolated localities there fide Kurek et al. 2017. For a map see Sindaco et al. 2013. 
EtymologyNamed after the length of this species which is unusually large for European snakes.

The genus was named after the Greek “za” = great, and “menos” = strength. 
References
  • Allentoft, M.E.; Rasmussen, A.R.; Kristensen, H.V. 2018. Centuries-Old DNA from an Extinct Population of Aesculapian Snake (Zamenis longissimus) Offers New Phylogeographic Insight. Diversity 10: 14, doi:10.3390/d10010014 - get paper here
  • Aßmann, Otto 2021. Man trifft sich an der Mauer. Elaphe 2021 (4): 38-40
  • Aßmann, Otto & Daniel Renner 2021. Schutz und Förderung der Äskulapnatter in Bayern. Elaphe 2021 (4): 42-49
  • Aßmann, Otto & Daniel Renner 2021. Zwischen Wildnis und Heizkeller – Aufenthaltsorte der Äskulapnatter in Landschaften, Lebensräumen und Habitaten an ihrer nordwestlichen Verbreitungsgrenze in Bayern. Elaphe 2021 (4): 34-37
  • Aßmann, Otto & Daniel Renner 2021. Grenzgänger: Neues zur Verbreitung der Äskulapnatter im bayerisch-österreichischen Grenzgebiet. Elaphe 2021 (4): 26-33
  • Bălescu, Carmen Daniela & Bușe-Dragomir Luminița 2023. PRELIMINARY OBSERVATIONS REGARDING THE HERPETOLOGICAL FAUNA FROM CORCOVA (MEHEDINȚI COUNTY). Analele Universitătii din Craiova, seria Agricultură - Montanologie - Cadastru (Annals of the University of Craiova - Agriculture, Montanology, Cadastre Series) 53 (2): 7-20
  • Berroneau, M. et al. 2010. Guide des Amphibiens et Reptiles d’Aquitaine. Association Cistude Nature, 180 pp.
  • Beshkov, V & Nanev, K 2006. The Amphibians and Reptiles in Bulgaria. Pensoft Series Faunistica 46, 120 pp.
  • Bezman-Moseyko, Olga S. 2010. New Records of Three Rare Snake Species from the Left Bank of Dniester River (Pridnestrovie). Russ. J. Herpetol. 17 (4): 280-284 - get paper here
  • Billy, G., Barbraud, C., Dahirel, M., & Bonnet, X. 2024. Does strict protection status harm snake populations in a temperate forest? Journal for Nature Conservation, 126683 - get paper here
  • Boettger, O. 1889. Herpetologische Miscellen. [I Epirus, II Corfu, III Kamerun, IV Gross-Namaland, V Transvaal, VI. Pondoland, VII Madagascar, VIII Madras, IX Java, X Nias, XI Nordwest-Peru]. Bericht senckenb. naturf. Ges. (Frankfurt): 267-316 - get paper here
  • Boettger, O. [as O. Böttger] 1880. Die Reptilien und Amphibien von Syrien, Palaestina und Cypern. Ber. Senckenb. Naturforsch. Ges., Frankfurt/M., 1879-1880: 132-219 - get paper here
  • Böhme, W. 1993. Äskulapnatter (Elaphe longissima LAURENTI, 1768). In: Böhme, W. (ed.), Handbuch der Reptilien und Amphibien Europas. Wiesbaden, Aula Verlag, pp. 331-372
  • Boulenger, G. A. 1913. The snakes of Europe. Methusen & Co. Ltd, London, 151 pp.
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Boyan Petrov; Pavel Stoev; Vladimir Beshkov 2001. A review on the species composition and distribution of the amphibians (Amphibia) and reptiles (Reptilia) in the Eastern Rhodopes Mountain, Bulgaria [in Russian] Hist. Nat. Bulgarica 13: 127-153 - get paper here
  • Bringsøe, H. 2015. Kæmpen vi ikke længere har plads til i Danmark – Iagttagelser af Æskulapsnog i Sydtyskland. Habitat 12: 6-17
  • Cabela, A., & Tiedemann, F. 1985. Atlas der Amphibien und Reptilien Osterreichs. Naturhist. Mus. in Wien, Neue Denkschriften Band 4 80 pp.
  • Canova,Luca & Manuela Marchesi 2007. Amphibian and reptile communities in eleven Sites of Community Importance (SCI): relations between SCI area, heterogeneity and richness. Acta Herpetologica 2 (2): 87-96
  • Capula, M.; L. Luiselli & L. Rugiero 2008. Variazioni della dieta in relazione allo stato di conservazione dei siti in Hierophis viridiflavus e Zamenis longissimus (Reptilia: Colubridae). Herpetologia Sardiniae (Claudia Corti, ed.), Edizioni Belvedere, 504 pp.
  • Christopoulos, A., Kotselis, C., Stefanopoulos, P., Zevgolis, Y.G. 2023. New distribution records for the Aesculapian snake Zamenis longissimus in Greek Thrace. Herpetological Bulletin, 164: 33–36. e_Zamenis_longissimus_in_Greek_Thrace - get paper here
  • CLEMENS, DAVID J. & STEVEN J. R. ALLAIN 2020. First record of the aesculapian snake (Zamenis longissimus) in South Wales. Herpetological Bulletin 152: - get paper here
  • Cogălniceanu, Dan; Laurentiu Rozylowicz, Paul Székely, Ciprian Samoilă, Florina Stănescu, Marian Tudor, Diana Székely, Ruben Iosif 2013. Diversity and distribution of reptiles in Romania. ZooKeys 341 (2013): 49-76 - get paper here
  • Covaciu-Marcov S.-D. et al. 2007. Contributions to the study of the geographical distribution of the herpetofauna in the North-East area of Arad County in Romania. Herpetologica Romanica 1: 62-69
  • Covaciu-Marcov S.-D.; Ghira, I.; Cicort-Lucaciu A.-St., Sas I.; Strugariu, A. & Bogdan H. V. 2006. Contributions to knowledge regarding the geographical distribution of the herpetofauna of Dobrudja, Romania. North-Western Journal of Zoology 2 (2): 88-125 - get paper here
  • Covaciu-Marcov, S.D., Cicort-Lucaciu, A.S., Dobre, F., Ferenţi, S., Birceanu, M., Mihuţ, R. & Strugariu, A. 2009. The herpetofauna of the Jiului Gorge National Park, Romania. North-Western Journal of Zoology 5 (Suppl. 1): 1-78
  • Covaciu-Marcov, S.D., Cicort-Lucaciu, A.S., Sas, I.; Mosu, A.G.; Toth, B. 2008. Contributions to the knowledge of the composition and geographical distribution of the Western Maramures County Herpetofauna. Herpetologica Romanica 2: 27-36
  • ÇAKMAK, Mücahit; Bahadır AKMAN and Mehmet Zülfü YILDIZ 2018. HERPETOFAUNA OF BARTIN PROVINCE (NORTHWEST BLACKSEA REGION, TURKEY). South Western Journal of Horticulture, Biology and Environment P-Issn: 2067- 9874, E-Issn: 2068-7958<br>Vol.8, No.2, 2017 pp.89-102 - get paper here
  • DI NICOLA, MATTEO RICCARDO 2019. A revised dichotomous key to the snakes of Italy (Reptilia, Squamata, Serpentes), according to recent systematic updates. Zootaxa 4686 (2): 294–296 - get paper here
  • Dossenbach, H.D., & Ganz, A. 1977. Die Reptilien der Schweiz. Naturf. Gesell. Schaffhausen, Switzerland, Neujahrsblatt nr. 30, 31 pp.
  • Dunaev E.A., Orlova V.F. 2003. Diversity of snakes (on the materials of the exposition of the Zoological Museum of the Moscow State University). Moscow: Moscow State University Press, 376 pp
  • Dunaev E.A., Orlova V.F. 2017. Amphibians and reptiles of Russia. Atlas and determination. 2nd. ed. Moscow: Phyton XXI, 328 p
  • Düşen, S.; İ.H. Uğurtaş & F.N. Altunel 2010. Nematode parasites of the smooth snake, Coronella austriaca Laurenti, 1768 and the Aesculapian snake, Zamenis longissimus (Laurenti, 1768) (Ophidia: Colubridae), collected from North-Western Turkey. North-Western Journal of Zoology 6 (1): 086-089
  • DYUGMEDZHIEV, ANGEL 2020. Tail vibration – a newly described defensive behaviour of the Aesculapian snake Zamenis longissimus. Herpetological Bulletin 154: - get paper here
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Esterbauer, H. 2014. Die Äskulapnatter, Zamenis longissimus (Laurenti, 1768) - Erstnachweis einer amelanistischen Farbmutation in Österreich. ÖKO·L 36/2: 7-11
  • Filippi, Ernesto 2003. The effects of timbering on a snake community of a Mediterranean area of central Italy. Amphibia-Reptilia 24 (1): 75-78 - get paper here
  • Geniez, Philippe 2018. Snakes of Europe, North Africa and the Middle East. Princeton University Press, 384 pp. - get paper here
  • Gezova, Simona; Peter Drugac, Adrian Purkart, Daniel Jablonski 2018. Albinism in Two Snake Species Recorded from Slovakia. Russ. J. Herpetol. 25 (1): 79-82 - get paper here
  • Golder, F. 1985. Ein gemeinsamer Massen-Eiablageplatz von Natrix natrix helvetica (LACEPEDE, 1789) und Elaphe longissima longissima (LAURENTI, 1768), mit Daten über Eizeitigung und Schlupf (Serpentes: Colubridae). Salamandra 21 (1): 10-16 - get paper here
  • Gomille, A. 2002. Die Äskulapnatter (Elaphe longissima) - Verbreitung und Lebensweise in Mitteleuropa. Edition Chimaira, Frankfurt, 158 pp.
  • Gruber, U. 2009. Die Schlangen Europas, 2. Aufl. Kosmos Naturführer, 266 pp.
  • Guiller, G. 2009. Déclin et biologie d’une population de Zamenis longissimus (Laurenti, 1768) (Serpentes, Colubridae) en Loire-Atlantique / Biological data and impact of village urbanization on Zamenis longissimus (Laurenti, 1768) (Ophidia, Colubridae) in Loire-Atlant Bull. Soc. Herp. France (132):85-114 - get paper here
  • GVOZDENOVIĆ, S., & IKOVIĆ, V. 2022. CHECKLIST OF AMPHIBIANS AND REPTILES OF MONTENEGRO AND THEIR CONSERVATION STATUS. Natura Montenegrina, Podgorica, 2022, 14 (2): 93-105 - get paper here
  • Gvozdenović, Slađana; Vuk Ikovic, Marko Nikolić 2021. Dicephalism in Aesculapian snake, Zamenis longissimus (Serpentes, Colubridae) from Montenegro. Herpetology Notes 14: 649-651
  • Hallmen, M. 2003. Freilandterrarien für Schlangen. Natur und Tier Verlag (Münster),157 pp. - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Heimes, P. 1994. Untersuchungen zur Ökologie und zum Verhalten der Äskulapnatter, Elaphe longissima (Laurenti, 1768) im Rheingau-Taunus. PhD Dissertation, University of Bonn, Germany, - get paper here
  • Heimes, P. & Waitzmann, M. 1993. Die Äskulapnatter (Elaphe longissima [Laurenti, 1768]) in Deutschland (Reptilia: Serpentes: Colubridae). Zool. Abh. Mus. Tierk. Dresden, 47 (2): 157-192 [1994]
  • Hoser, R. 2012. A taxonomic revision of the Colubrinae genera Zamenis and Orthriophis with the creation of two new genera (Serpentes: Colubridae). Australasian J. Herpetol. 11: 59-64 - get paper here
  • Jaeschke, J. 1971. Zur Einbürgerung der Äskulapnatter in Oberhessen. Salamandra 7 (2): 85 - get paper here
  • Joger, U.; Fritz, U.; Guicking, D.; Kalyabina-Hauf, S.; Nagy, Z.T. & Wink, M. 2007. Phylogeography of western Palaearctic reptiles – Spatial and temporal speciation patterns. Zoologischer Anzeiger 246: 293–313 - get paper here
  • Kabisch, K. & Engelhard, H. 1964. Beobachtungen an Reptilien in der Umgebung von Varna-Goldsand. Aquarien Terrarien 11 (6): 194-197
  • Kaiser, H.; Crother, B.I.; Kelly, C.M.R.; Luiselli, L.; O’Shea, M.; Ota, H.; Passos, P.; Schleip, W.D. & Wüster, W. 2013. Best Practices: In the 21st Century, Taxonomic Decisions in Herpetology are Acceptable Only When Supported by a Body of Evidence and Published via Peer-Review. Herpetological Review 44 (1): 8-23
  • Kamali, Kamran 2020. A Field Guide to the Reptiles and Amphibians of Iran. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 574 pp.
  • KAMMEL, W. 2007. Aktivität und Nahrungserwerb der Äskulapnatter, Zamenis longissimus longissimus (LAURENTI, 1768) in Österreich. Herpetozoa 20 - get paper here
  • Kammel, Werner 2009. External morphology and sex ratio of the Aesculapian Snake Zamenis longissimus longissimus (Laurenti, 1768), in Austria (Squamata: Serpentes: Colubridae). Herpetozoa 21 (3/4):99-121 - get paper here
  • KANUCH, P. & BALÁZ, P. 2005. Bat as a prey of Elaphe longissima (LAURENTI, 1768) [Short Note]. Herpetozoa 18: 92- 93 - get paper here
  • Katarzyna Kurek, Kamil Najberek, Bartłomiej Zając, Stanisław Bury, Adam M. Ćmiel, Grzegorz Baś, and Bartłomiej Najbar 2017. Changes in Distribution of Aesculapian Snake and Implications for Its Active Conservation in Poland. Polish Journal of Ecology 65 (4): 422-431 - get paper here
  • KAYA, Nilgün; Oya ÖZULUĞ 2017. THE HERPETOLOGICAL COLLECTION OF ZOOLOGY MUSEUM, ISTANBUL UNIVERSITY. Anadolu Üniversitesi Bilim ve Teknoloji Dergisi C- Yaşam Bilimleri ve Biyoteknoloji<br>(Anadolu University Journal of Science and Technology C- Life Sciences and Biotechnology) - get paper here
  • Khrabrov I. E., Sadykov R. K., Shapovalov M. I. 2022. New findings of the Aesculapian skidder Zamenis longissimus (Laurenti, 1768) (Colubridae, Reptilia) in the Republic of Adygea. [in Russian] Current Studies in Herpetology 22 (1-2): 73 - get paper here
  • Kolodziej, Kai 2017. Urbanes Herping – Reptilien und Amphibien um eine Millionenstadt. Terraria-Elaphe 2017 (4): 44-49 - get paper here
  • König, D. 1985. Langjährige Beobachtungen an der Äskulapnatter Elaphe longissima (LAURENTI 1768). Salamandra 21 (1): 17-39 - get paper here
  • Kovar, Roman; Marek Brabec , Radovan Vita , Roman Vodicka & Vlastimil Bogdan 2016. Nesting and over-wintering sites of Aesculapian snake, Zamenis longissimus, in an anthropogenic landscape in the northern extreme of its range. Herpetological Bulletin (136) - get paper here
  • Kovar, Roman; Marek Brabec, Radovan Vita, and Radomir Bocek 2014. Mortality Rate and Activity Patterns of an Aesculapian Snake (Zamenis longissimus) Population Divided by a Busy Road. Journal of Herpetology Mar 2014, Vol. 48, No. 1: 24-33. - get paper here
  • Kovar, Roman; Marek Brabec, Radovan Vita, Roman Vodicka & Vlastimil Bogdan 2016. Habitat use of the Aesculapian Snake, Zamenis longissimus, at the northern extreme of its range in northwest Bohemia. Herpetological Bulletin (136) - get paper here
  • Kreiner, G. 2009. Die Schlangen Nordspaniens - Reiseeindrücke aus dem Kantabrischen Gebirge und angrenzenden Gebieten. Draco 10 (39): 38-48 - get paper here
  • Krizmanić, I. 2018. CONTRIBUTION TO THE HERPETOFAUNA OF SERBIA-DISTRIBUTION OF REPTILES IN KOSOVO AND METOHIJA PROVINCE. Bulletin of Natural Sciences Research 9 (2): - get paper here
  • Krofel, M. 2004. First record of albino Aesculapian Snake (Elaphe longissima) in Slovenia. Natura Sloveniae 6 (2): 53-56
  • Kühnel, Klaus-Detlef; Arno Geuger & Richard Podloucky 2012. Die Rote Liste der Amphibien und Reptilien Deutschlands. Teil 1: Werdegang und Ergebnisse. Terraria-Elaphe 2012 (2): 92-100 - get paper here
  • Kwet, A. 2007. Zamenis longissimus (Laurenti, 1768) Äskulapnatter. Reptilia (Münster) 12 (67): 51-54 - get paper here
  • Kwet, A. 2010. Reptilien und Amphibien Europas. Kosmos-Verlag, Stuttgart, 252 pp.
  • Kwet, Axel & Benny Trapp 2014. Liste der Reptilien Europas. Draco 15 (60): 72-79 - get paper here
  • Lacepède, B. G. E. 1803. Histoire Naturelle des Quadrupèdes Ovipares et de Serpens. Vol. 4. Paris: Chez Saugrain, 386 pp. - get paper here
  • Laufer, H.; Fritz, C. & Sowig, P. (eds.) 2007. Die Amphibien und Reptilien Baden-Württembergs. Ulmer Verlag, Stuttgart, 807 pp.
  • Laurenti, J. N. 1768. Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austracorum, quod authoritate et consensu. Vienna, Joan. Thomae, 217 pp. - get paper here
  • Lelièvre, Herve; Coraline Moreau, Gabriel Blouin-Demers, Xavier Bonnet, and Olivier Lourdais 2012. Two Syntopic Colubrid Snakes Differ In Their Energetic Requirements and In Their Use of Space. Herpetologica 68 (3): 358-364. - get paper here
  • Lelièvre, Hervé; Legagneux, Pierre; Blouin-Demers, Gabriel; Bonnet, Xavier; Lourdais, Olivier 2012. Trophic niche overlap in two syntopic colubrid snakes (Hierophis viridiflavus and Zamenis longissimus) with contrasted lifestyles. Amphibia-Reptilia 33 (1): 37-44 - get paper here
  • Lenk, P.; Joger, U. & Wink, M. 2001. Phylogenetic relationships among European ratsnakes of the genus Elaphe Fitzinger based on mitochondrial DNA sequence comparisons. Amphibia-Reptilia 22 (3): 329-339 - get paper here
  • Leptien, R. 2011. Tiere unserer Heimat: Die Äskulapnatter. Reptilia (Münster) 16 (89): 79-82 - get paper here
  • Lotze, H. U. 1975. Zum Paarungsverhalten der Äskulapnatter, Elaphe longissima. Salamandra 11 (2): 67-76 - get paper here
  • Major, T. 2024. The ecology, biogeography, and taxonomy of isolated snake populations. PhD thesis, Bangor University - get paper here
  • Major, Tom; Devlan R. Alkins, Lauren Jeffrey & Wolfgang Wüster 2020. Marking the un-markable: visible implant elastomer in wild juvenile snakes. The Herpetological Journal 30: 173-176 - get paper here
  • Molen, Sjef van der 2006. A two-headed Zamenis longissimus Podarcis 7 (1/2): 7-8 - get paper here
  • Naulleau, G. 1987. Les serpents de France, 2.ed. Rev. Francaise d'Aquariologie, Herpet. (Nancy) 56 pp. - get paper here
  • NAUMOV, BORISLAV; NIKOLAY TZANKOV, KRASIMIR DONCHEV, BOYAN PETROV, ANDREI STOJANOV, GEORGI POPGEORGIEV, IVELIN MOLLOV, VLADIMIR BESHKOV 2016. The Herpetofauna (Amphibia and Reptilia) of Vrachanska Planina Mountains - Species Composition, Distribution and Conservation. In: Bechev, D. & Georgiev, D. (Eds.), Faunistic diversity of Vrachanski Balkan Nature Park. ZooNotes, Supplement 3, Plovdiv University Press, Plovdiv - get paper here
  • Nečas, Petr; Modry, David & Zavadil, Vít 1997. Czech Recent and Fossil Amphibians and Reptiles. Edition Chimaira, Frankfurt, 94 pp.
  • Niebergall, P. 2003. Katarakt (Grauer Star) bei einer Äskulapnatter, Elaphe longissima (LAURENTI 1768). Sauria 25 (3): 29-31 - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Petrov B. P., N. Tzankov, H. Strijbosch, G. Popgeorgiev, V. Beshkov 2006. The herpetofauna (Amphibia and Reptilia) of the Western Rhodopes mountain (Bulgaria and Greece). In: Beron P. (ed.). Biodiversity of Bulgaria. 3. Biodiversity of Western Rhodopes (Bulgaria and Greece) I. Pensoft & Nat. Mus. Natur. Hist., Sofia, pp. 863-912
  • Polovic, Lidija and Natalija Cadenovic. 2014. The herpetofauna of Ljubisnja Mountain, Montenegro. Herpetozoa 26 (3/4): 193-194 - get paper here
  • Pottier G., Paumier J.-M., Tessier M., Barascud Y., Talhoët S., Liozon R., D’Andurain P., Vacher J.-P., Barthe L., Heaulmé V., Esslinger M., Arthur C.-P., Calvet A., Maurel C. & Redon H. 2008. Atlas de répartition des reptiles et amphibiens de Midi-Pyrénées. Les atlas naturalistes de Midi-Pyrénées. Nature Midi-Pyrénées, Toulouse, 126 pp.
  • Rajabizadeh, M. 2018. Snakes of Iran. [in Farsi] Tehran: Iranshenasi, 496 pp. [2017 ?] - get paper here
  • RASTEGAR-POUYANI, NASRULLAH; HAJI GHOLI KAMI, MEHDI RAJABZADEH, SOHEILA SHAFIEI AND STEVEN CLEMENT ANDERSON 2008. Annotated Checklist of Amphibians and Reptiles of Iran. Iranian Journal of Animal Biosystematics 4 (1): 7-30
  • Reading, C. J.; L. M. Luiselli, G. C. Akani, X. Bonnet, G. Amori, J. M. Ballouard, E. Filippi, G. Naulleau, D. Pearson, L. Rugiero 2010. Are snake populations in widespread decline? Biol. Lett. 2010 6 777-780; DOI: 10.1098/rsbl.2010.0373 - get paper here
  • Renner, Daniel & Otto Aßmann 2017. Die Schlingnatter als Prädator der Äskulapnatter. Terraria-Elaphe 2017 (6): 72 - get paper here
  • Rödel, M.-O. & R. BUSSMANN 1990. Die Amphibien und Reptilien der Julischen Alpen. Sauria 12 (4): 15-21 - get paper here
  • Rugiero, Lorenzo; Luca Luiselli, Massimo Capula, Fabio Petrozzi, Massimiliano Di Vittorio, Nic Pacini, Leonardo Vignoli, Daniele Dendi, Giuliano Milana, Giovanni Amori & Roger Meek 2020. Snakes and the Eternal City: variation in community metrics, body size and population density of snakes in contrasted landscapes of suburban Rome (Italy). The Herpetological Journal 30: 69-82 - get paper here
  • Rugiero, Lorenzo; Massimo Capula, Dario Capizzi, Giovanni Amori, Giuliano Milana, Manuela Lai, Luca M. Luiselli 2018. Long-term observations on the number of roadkilled Zamenis longissimus (LAURENTI, 1768) in a hilly area of Central italy. Herpetozoa 30 (3/4): 212 - 217 - get paper here
  • Salvi D, Mendes J, Carranza S, Harris DJ. 2018. Evolution, biogeography and systematics of the western Palaearctic Zamenis ratsnakes. Zoologica Scripta 47 (4): 441-461 - get paper here
  • Santos, X., Pleguezuelos, J. M., Chergui, B., Geniez, P., & Cheylan, M. 2022. Citizen-science data shows long-term decline of snakes in southwestern Europe. Biodiversity and Conservation, 1-17 - get paper here
  • Schlüter, U. 2006. Die Herpetofauna der bulgarischen Schwarzmeerküste - Teil 3: Schlangen. Elaphe 14 (2): 59-66
  • Schlüter, U. 2009. Die Schlangenfauna Europas. Draco 10 (39): 4-21 - get paper here
  • Schmidt B, Kranželić D, Majetić Đ, Lauš B, Štih A, Koren T 2020. Distribution and conservation status of the herpetofauna of Dugi Otok Island, Croatia. Herpetozoa 33: 191-205 - get paper here
  • Schmidt, K. E, & INGER, R. F. 1957. Living Reptiles of the World. Doubleday & Co., 287 pp. - get paper here
  • Schulz, K. D. 1996. Eine Monographie der Schlangengattung Elaphe Fitzinger. Bushmaster, Berg (CH): 1-460
  • Schulz, Klaus-Dieter 1996. A monograph of the colubrid snakes of the genus Elaphe Fitzinger. Koeltz Scientific Books, 439 pp.
  • Schweiger, M. 2012. Die Schlangenfauna der kroatischen Insel Krk. Ophidia 6 (2): 2-18 - get paper here
  • Schweiger, Mario 1994. Erstnachweis von Elaphe longissima (Laurenti, 1768) für die zentrale Osttürkei (Squamata: Serpentes: Colubridae). Herpetozoa 7 (3/4): 149-151 - get paper here
  • Sclater, W. L. 1891. List of snakes in the Indian Museum. Trustees of the Indian Museum, Calcutta, 79 pp. - get paper here
  • Sehnal, P. & Schuster, A. 1999. Herpetologische Beobachtungen auf der Kvarnerinsel Cres, Kroatien. Ergebnisse von fünf Exkursionen. Herpetozoa 12 (3-4): 163-178 - get paper here
  • Sindaco, R.; Alberto Venchi & Cristina Grieco 2013. The Reptiles of the Western Palearctic, Volume 2: Annotated Checklist and Distributional Atlas of the Snakes of Europe, North Africa, Middle East and Central Asia, with an Update to Volume 1. Edizioni Belvedere, Latina (Italy), 543 pp. - get paper here
  • Sindaco, Roberto; Alberto Venchi, Giuseppe M. Carpaneto, Marco A. Bologna 2000. The Reptiles of Anatolia: a Checklist and Zoogeographical analysis. Biogeographia, 21-2000: 441-554. - get paper here
  • Skubowius, Bernd 2008. Das Erlebnis Schlangenprozession von Cocullo. Ophidia 2 (2): 2-10 - get paper here
  • Sobolenko, L.Yu., Nekrasova, O.D., Sorokina, S.I., Moroz, L.M. 2018. Current state of reptiles of western podillia. Ukrainian Journal of Ecology, 8(1): 706–711 - get paper here
  • Sos, T. 2008. Review of recent taxonomic and nomenclatural changes in European Amphibia and Reptilia related to Romanian herpetofauna. Herpetologica Romanica 2: 61-91
  • Stevens, Kevin 1995. The European ratsnakes of the genus Elaphe. British Herpetological Society Bulletin (54): 10-20
  • Stille, Bo and Marie Stille 2017. The Herpetofauna of Corfu and Adjacent Islands. Edition Chimaira, Frankfurt am Main, Germany (www.chimaira.de). 354 pp.,
  • Stoev, Pavel 2000. On the distribution, biology and ecology of amphibians and reptiles in the Derventski Heights and the Sakar Mountain, South-East Bulgaria Hist. Nat. Bulgarica 12: 59–69 - get paper here
  • STRACHINIS, ILIAS. 2021. First evidence on the occurence of Greek Newt Lissotriton graecus and the Aesculapian snake Zamenis longissimus on Evia Island, Greece - Parnassiana Archives 9: 115-117. Parnassiana Archives 9: 115–117. - get paper here
  • Strödicke, Martin and Birgit Gerisch. 1999. Morphologiasche Merkmalsvariabilität bei Elaphe longissima (Laurenti, 1768), unter besonderer Berücksichtigung zweier isolierter Populationen an der Nordgrenze des Artareals (Squamata: Serpentes: Colubridae). Herpetozoa 11 (3/4):121-139 [1998] - get paper here
  • Strugariu, A., Zamfirescu, A.R., Nicoară, A., Gherghel, I., Sas, I., Puşcaşu, C.M. & Bugeac, T. 2008. Preliminary data regarding the distribution and status of the herpetofauna in Iaşi County (Romania). North-Western Journal of Zoology 4 (Suppl.1): S1-S23 - get paper here
  • Strugariu, A.; Sahlean, T.C.; Volosciuc-Hutuleac, M.V. & Puscasu, C.M. 2006. Preliminary data regarding the distribution of reptilian fauna in Suceava County (Romania). North-Western Journal of Zoology 2 (1): 39-43 - get paper here
  • Tóth, Tamás, László Krecsák, Thomas Madsen and Beáta Újvári 2002. Herpetofaunal locality records on the Greek Islands of Corfu (Amphibia, Reptilia). Herpetozoa 15 (3/4):149-169 - get paper here
  • Trapp, B. 2007. Reptilien und Amphibien des griechischen Festlandes. Natur und Tier Verlag (Münster), 300 pp. - get paper here
  • Trutnau, L. 1975. Europäische Amphibien und Reptilien. Belser Verlag, Stuttgart 212 pp.
  • Tsellarius A. Y. 2021. Specific composition, numbers and biotopical distribution of snakes (Serpentes) on the southern slope of the Navagir mountain ridge (Abrau Peninsula) and some problems of its preservation [in Russian]. Current Studies in Herpetology 21 (3-4): 144 - get paper here
  • Tuniyev B.S., Tuniyev S.B., Avcı A., Ilgaz Ç. 2014. Herpetological studies in Eastern and North-Eastern Turkey. Current Studies in Herpetology 14 (1/2): 44–53. Резюме - get paper here
  • Tuniyev, B.S.; N.L. Orlov, N.B. Ananjeva and A.L. Aghasyan 2019. Snakes of the Caucasus: taxonomic diversity, distribution, conservation. St. Petersburg, Moscow. KMK Scientific Press. 2019. 276 pp. - get paper here
  • Uhrin, Marcel; Peter Havaš, Martin Minařík, Karel Kodejš, Imrich Bugoš, Stanislav Danko, Tomáš Husák, Daniel Koleska and Daniel Jablonski 2016. Distribution updates to amphibian and reptile fauna for the Republic of Macedonia. Herpetology Notes 9: 201-220 - get paper here
  • Utiger, Urs, Notker Helfenberger, Beat Schätti, Catherine Schmidt, Markus Ruf and Vincent Ziswiler 2002. Molecular systematics and phylogeny of Old World and New World ratsnakes, Elaphe Auct., and related genera (Reptilia, Squamata, Colubridae). Russ. J. Herpetol. 9 (2): 105-124. - get paper here
  • Vaccaro, Angelo & Turrisi, G.F. 2007. Ritrovamento di Zamenis longissimus (Laurenti, 1768) (Reptilia, Colubridae) sull’Isola d’Elba (Toscana, Italia). Acta Herpetologica 2 (1): 59-63
  • Veith, G. 1991. Die Reptilien Bosniens und der Herzegowina. Teil 2. Herpetozoa 4 (1-2): 1-96 - get paper here
  • Venchi, Alberto and Roberto Sindaco 2006. Annotated checklist of the reptiles of the Mediterranean countries, with keys to species identification. Part 2 -Snakes (Reptilia, Serpentes). Annali del Museo Civico di Storia Naturale "G. Doria", Genova, XCVIII: 259-364
  • Waitzmann, Michael 1993. Zur Situation der Äskulapnatter Elaphe longissima (Laurenti, 1768) in der Bundesrepublik Deutschland. Mertensiella (3): 115-134
  • Waitzmann, Michael; Annette Zitzmann, Andreas Malten & Otto Aßmann 2021. Die Äskulapnatter in Deutschland. Elaphe 2021 (4): 12-17
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Weinbeer, Niko; Volker Harport & Harald Löffler 2017. Die Äskulapnatter – auf der Suche nach einem sagenumwobenen Reptil in heimischen Gefilden. Terraria-Elaphe 2017 (3): 46-49 - get paper here
  • Wilson, Matt J., Bo Stille and Marie Stille. 2014. Herpetofauna of Paxos, Ionian Islands, Greece, including two species new to the island. Herpetozoa 27 (1/2): 108-112 - get paper here
  • Wirth, M. 2009. Königinnen der europäischen Schlangen: Vierstreifennatter - Elaphe quatuorlineata), Äskulapnatter (Zamenis longissimus) und Leopardnatter (Zamenis situla). Draco 10 (39): 61-76 - get paper here
  • Wirth, Michael 2013. Das Geheimnis der Feuerschneisen. Terraria-Elaphe 2013 (3): 36-43 - get paper here
  • Wütschert, R. 1984. Neues uber die Reptilienfauna der Insel Korfu. Salamandra 20 (4): 221-228 - get paper here
  • Žagar, A., Carretero, M. A., Krofel, M., Lužnik, M., Podnar, M. & Tvrtković, N. 2014. Reptile survey in Dinara Mountain (Croatia) revealed the southernmost known population of Horvath’s rock lizard (Iberolacerta horvathi). Nat. Croat. 23 (1): 235–240 - get paper here
  • Zang, R. 1904. Über horizontale und vertikale Verbreitung einiger reptilien und Amphibien Deutschlands und der Schweiz. Zool. Anz. 28: 249-251 - get paper here
  • ZAUNER, J. 2021. Auf Reptiliensuche im Soca-Tal (Slowenien). Sauria 43 (1): 2-8 - get paper here
  • Zitzmann, Annette; Andreas Malten, Jenny Behm & Michael Waitzmann 2021. Schutz und Förderung der Äskulapnatter in Hessen und Baden-Württemberg. Elaphe 2021 (4): 18-25
 
External links  
Is it interesting? Share with others:


Please submit feedback about this entry to the curator