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Andinosaura petrorum (KIZIRIAN, 1996)

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymProctoporus petrorum KIZIRIAN 1996: 122
Proctoporus petrorum — DOAN & SCHARGEL 2003
Riama petrorum — DOAN & CASTOE 2005
Andinosaura petrorum — SÁNCHEZ-PACHECO et al. 2017 
DistributionEcuador (Morona-Santiago), elevation 2195 m

Type locality: trail between Sevilla de Oro and Méndez, on the eastern slopes of the mountains between Cerro Negro and Pailas, Morona-Santiago, Ecuador.  
Reproductionoviparous 
TypesHolotype: USNM 196267, adult male 
DiagnosisDiagnosis: -(1) Frontonasal slightly shorter than frontal; (2) nasoloreal suture absent; (3) supraoculars three or four; second, third, and fourth, or first, second, and third in contact with ciliaries; (4) superciliary series incomplete, one; (5) supralabial-subocular fusion absent; (6) postoculars two or three; (7) postparietals two; (8) supratympanic temporals three; (9) genials two, transverse sutures not perpendicular with respect to midline of body; (10) dorsals rectangular, juxtaposed, heavily striated/keeled; (11) longitudinal dorsal scale rows in male 21, in females 21 or 22; (12) transverse dorsal scale rows in male 33, in females 31-33; (13) transverse ventral scale rows in male 19, in females 19-22; (14) lateral scale rows two or three; (15) femoral pores in male 4/5, in females zero or four; scales between femoral pores eight; (16) subdigital scales on Toe I five or six; (17) limbs not overlapping when adpressed against body in adults; (18) anterior cloacal plate scales paired; (19) hemipenial morphology unknown; (20) dorsum olive to dark brown, large ocelli present laterally; venter unicolored olive or dark brown with cream along scale many sutures, especially posterior and longitudinal sutures on belly; subcaudally nearly unicolored dark brown, or olive.
Proctoporus petrorum can be distinguished by the following (condition for P. petrorum in parentheses): All Ecuadorian congeners (except P. hypostictus): femoral pores in males five or more (four). All Ecuadorian congeners except P. anatoloros and P. vespertinus: scales between femoral pores six or fewer (eight). P. hypostictus: genials three (two). P. vespertinus: anterior superciliaries two (one). P. luctuosus group: lateral scales in wide band, more than two (two or three rows of granular scales), superciliary series complete (incomplete), total femoral pores in males 15 or more (nine). P. achlyens, P. laevis, P. luctuosus: nasoloreal suture complete (absent). P. columbianus: dorsal scales smooth (keeled), nasoloreal suture complete (absent). P. striatus: large lateral ocelli absent (present); superciliary series complete (incomplete) limbs overlapping (not overlapping) when adpressed against body. (Kizirian 1996)


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Comment 
EtymologyThe specific epithet is a Latin name, petrus, -i, in the genitive plural case, that translates to "of the Peters." The name honors Wilhelm Peters, in recognition of his contributions to gymnophthalmid systematics (see in particular his monograph [Peters,1862]), and the collectors of the type material, Peter D. Spoecker and James A. Peters, the latter of whom made significant contributions to the understanding of the Ecuadorian herpetofauna and made many collections of Proctoporus. 
References
  • Arteaga, A.; Bustamante, L.; Vieira, J. 2024. Reptiles of Ecuador. Khamai Foundation & Tropical Herping, 1073 pp. - get paper here
  • Doan, T. M. & Castoe, T.A. 2005. Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society, 143: 405–416. - get paper here
  • Doan, T.M. & Schargel, W.E. 2003. Bridging the gap in Proctoporus distribution: a new species (Squamata: Gymnophthalmidae) from the Andes of Venezuela. Herpetologica 59 (1): 68-75 - get paper here
  • Kizirian, D. A. 1996. A review of Ecuadorian Proctoporus (Squamata: Gymnophthalmidae) with descriptions of nine new species. Herpetological Monographs 10: 85-155 - get paper here
  • Sánchez-Pacheco, S. , Aguirre-Peñafiel, V. , Torres-Carvajal, O. 2012. Lizards of the genus Riama (Squamata: Gymnophthalmidae): the diversity in southern Ecuador revisited. South American J. Herp. 7 (3): 259-275 - get paper here
  • Sánchez-Pacheco, S. J., Torres-Carvajal, O., Aguirre-Peñafiel, V., Sales-Nunes, P. M., Verrastro, L., Rivas, G. A., Rodrigues, M. T., Grant, T. and Murphy, R. W. 2017. Phylogeny of Riama (Squamata: Gymnophthalmidae), impact of phenotypic evidence on molecular datasets, and the origin of the Sierra Nevada de Santa Marta endemic fauna. Cladistics, doi:10.1111/cla.12203 [print: 2018] - get paper here
  • Sánchez-Pacheco, Santiago J.; David A. Kizirian, and Pedro M. Sales-Nunes 2011. A New Species of Riama from Ecuador Previously Referred to as Riama hyposticta (Boulenger, 1902) (Squamata: Gymnophthalmidae). American Museum Novitates (3719): 1-15 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
 
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