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Andinosaura vieta (KIZIRIAN, 1996)

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Higher TaxaGymnophthalmidae (Cercosaurinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common Names 
SynonymProctoporus vietus KIZIRIAN 1996: 145
Proctoporus vietus — TORRES-CARVAJAL 2001
Proctoporus vietus — DOAN & SCHARGEL 2003
Riama vieta — DOAN & CASTOE 2005
Andinosaura vieta — SÁNCHEZ-PACHECO et al. 2017 
DistributionEcuador (Cotopaxi)

Type locality: San Francisco de Las Pampas, Cotopaxi, Ecuador, 78° 58’ W, 00° 25’ S, 1800 m elevation.  
Reproductionoviparous 
TypesHolotype: MHNG 2437.48, adult male 
DiagnosisDiagnosis: (1) Frontonasal equal to or slightly shorter than frontal; (2) nasoloreal suture usually present; (3) supraoculars three or four, usually four, none in contact with ciliaries; (4) superciliary series complete, usually four; (5) supralabial-subocular fusion absent; (6) postoculars two or three, usually two; (7) postparietals two; (8) supratympanic temporals two or three, usually three; (9) genials three, transverse sutures perpendicular with respect to midline of body; (10) dorsals rectangular, juxtaposed, keeled, rugose; (11) longitudinal dorsal scale rows in males 14-18, in females 13-21; (12) transverse dorsal scale rows 29-32; (13) transverse ventral scale rows 20-23; (14) lateral scale rows 3-6; (15) femoral pores in males 8-10, in females 0-2, scales between femoral pores in males zero or one, usually zero, in females two; (16) subdigital scales on Toe I 6-9; (17) limbs overlapping when adpressed in adults; (18) anterior cloacal plate scales paired or unpaired; (19) hemipenial morphology unknown; (20) dorsum brown with faint or broken dorsolateral line, most distinct anteriorly, and above hind limb and on tail, sometimes distinct anterior to eye; large black spots sometimes present anterodorsally; small ocelli sometimes present laterally; venter black with white on transverse sutures or white with small black spots, especially laterally. Proctoporus vietus is distinguished from its congeners, except some P. striatus, by the presence of rugose scales on the dorsum and limbs. Proctoporus striatus has genials three (usually two); lateral scales in 0-3 (3-6) rows. (Kizirian 1996)


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Comment 
EtymologyThe specific epithet, vietus, is a Latin adjective that means wrinkled or shriveled and refers to the rugose scale surfaces, especially of the dorsal scales. 
References
  • Arteaga, A.; Bustamante, L.; Vieira, J. 2024. Reptiles of Ecuador. Khamai Foundation & Tropical Herping, 1073 pp. - get paper here
  • Doan, T. M. & Castoe, T.A. 2005. Phylogenetic taxonomy of the Cercosaurini (Squamata: Gymnophthalmidae), with new genera for species of Neusticurus and Proctoporus. Zoological Journal of the Linnean Society, 143: 405–416. - get paper here
  • Doan, T.M. & Schargel, W.E. 2003. Bridging the gap in Proctoporus distribution: a new species (Squamata: Gymnophthalmidae) from the Andes of Venezuela. Herpetologica 59 (1): 68-75 - get paper here
  • Kizirian, D. A. 1996. A review of Ecuadorian Proctoporus (Squamata: Gymnophthalmidae) with descriptions of nine new species. Herpetological Monographs 10: 85-155 - get paper here
  • Sánchez-Pacheco, S. J., Torres-Carvajal, O., Aguirre-Peñafiel, V., Sales-Nunes, P. M., Verrastro, L., Rivas, G. A., Rodrigues, M. T., Grant, T. and Murphy, R. W. 2017. Phylogeny of Riama (Squamata: Gymnophthalmidae), impact of phenotypic evidence on molecular datasets, and the origin of the Sierra Nevada de Santa Marta endemic fauna. Cladistics, doi:10.1111/cla.12203 [print: 2018] - get paper here
  • Sánchez-Pacheco, Santiago J.; David A. Kizirian, and Pedro M. Sales-Nunes 2011. A New Species of Riama from Ecuador Previously Referred to as Riama hyposticta (Boulenger, 1902) (Squamata: Gymnophthalmidae). American Museum Novitates (3719): 1-15 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Torres-Carvajal, O. 2001. Lizards of Ecuador: Checklist, Distribution, and Systematic References. Smithsonian Herp. Inf. Serv. (131): 1-35 - get paper here
 
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