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Anolis festae PERACCA, 1904

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Higher TaxaAnolidae, Iguania, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Veronica's Anole 
SynonymAnolis festae PERACCA 1904: 4
Anolis nigrolineatus WILLIAMS 1965
Anolis nigrolineatus — PETERS & DONOSO-BARROS 1970: 61
Anolis festae – PETERS & DONOSO-BARROS 1970: 53
Anolis nigrolineatus — WILLIAMS 1974
Dactyloa nigrolineata — NICHOLSON et al. 2012
Dactyloa festae — NICHOLSON et al. 2012
Dactyloa festae — NICHOLSON et al. 2018
Dactyloa nigrolineata — NICHOLSON et al. 2018
Anolis festae — POE 2025 
DistributionW Ecuador (lowlands), Colombia (Nariño)

Type locality: Balzar, Ecuador

nigrolineatus (invalid): Ecuador; Type locality: Machala, Provincia El Oro, Ecuador (exact locality in doubt, fide Williams 1965)  
Reproductionoviparous 
TypesHolotype: MRSN (= MSNTO) R2895; syntype: MSNG 37700
Holotype: MCZ 38940 [nigrolineatus] 
DiagnosisDiagnosis (nigrolineatus): Similar to A. punctatus Daudin but differing in color (lighter and with a black midvertebral line and faint dark markings on flanks and limbs) and in squamation (fewer scales in contact with the rostral, fewer lamellae under fourth toe). (Williams 1965)


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CommentSpecies groups: Dactyloa punctata species group (fide NICHOLSON et al. 2012).

RDB speciesID: 10899 [nigrolineatus]

Etymology (nigrolineatus): Named after Latin nigro, black, dark and Latin lineatus, lined. [“...A black middorsal line two scales wide...”]. (from Esteban Lavilla, pers. comm., May 2024) 
EtymologyNamed after Dr. Enrico Festa (1868-1939), an Italian zoologist. He visited Egypt, Palestine, Jordan, Lebanon, and Syria (1893) and collected in Panama and Ecuador (1895-1898). He worked for Museo e Instituto di Zoologia Sistematica dell'Universita di Torino (1899-1923) as Deputy Assistant Professor (1899), retiring as Honorary Vice Director (Beolens et al. 2011: 89). 
References
  • Arteaga, A.; Bustamante, L.; Vieira, J. 2024. Reptiles of Ecuador. Khamai Foundation & Tropical Herping, 1073 pp. - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Capocaccia, L. 1961. Catalogo dei tipi di Rettili del Museo Civico di Storia Naturale di Genova [MSNG]. Ann. Mus. Civ. Stor. Nat. Giacomo Doria 72: 86-111 - get paper here
  • Cruz-García K, Barreno M, Cuadrado S, Moretta-Urdiales M, Molina-Moreira N, Salas JA, Torres-Domínguez Á, Narváez AE 2023. Amphibians and reptiles of Isla Santay (Durán, Ecuador). Check List 19(3): 347-369 - get paper here
  • Cruz-García, K., N.Zapata-Salvatierra, A. E. Narváez, and J.Pérez-Correa 2026. Urban–Wild Interface Diversity: A Comprehensive Checklist of Herpetofauna of Guayaquil, Ecuador. Ecology and Evolution16, no. 5: e73504 - get paper here
  • Cuadrado, S. S., Loor, Y. A., Zambrano, R. H., & Narváez, A. E. 2023. Species Composition and Ecology of Diurnal Herpetofauna in Agroecosystems of the Ecuadorian Chocó. South American Journal of Herpetology, 29(1), 66-76 - get paper here
  • Medina-Rangel, Guido F.;, María Alejandra Pinto-E. and Martha L. Calderón-Espinosa 2018. First record of the Veronica´s Anolis Anolis festae (Squamata, Dactyloidae) in Colombia. Herpetology Notes 11: 725-728 - get paper here
  • Miyata, Kenneth I. 2013. Studies on the Ecology and Population Biology of Little Known Ecuadorian Anoles. Bull. Mus. Comp. Zool. Harvard 161(2):45-78 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2018. Translating a clade based classification into one that is valid under the international code of zoological nomenclature: the case of the lizards of the family Dactyloidae (Order Squamata). Zootaxa 4461 (4): 573–586 - get paper here
  • Peracca, M. G. 1904. Viaggio del Dr. Enrico Festa nell Ecuador e regioni vicine. Boll. Mus. Zool. Comp. Anat. Univ. Torino 19 (465): 1-41 - get paper here
  • Peters, James A. & Donoso-Barros, Roberto 1970. Catalogue of the Neotropical Squamata: Part II. Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 293 pp. - get paper here
  • Pinto-Erazo, M.A., Calderón-Espinosa, M.L., Medina-Rangel, G.F. & Méndez-Galeano, M.A. 2020. Herpetofauna from two municipalities of southwestern Colombia. Biota Colombiana 21(1): 41-57. DOI: 10.21068/c2020.v21n01a04 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Poe, Steven 2025. A Guide to the Anolis Lizards (Anoles) of Mainland Central and South America. Princeton University Press, 432 pp. - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Williams,E.E. 1965. South American Anolis (Sauria: Iguanidae): Two new species of the punctatus group. Breviora (233): 1-15 - get paper here
  • Williams,E.E. 1974. South American Anolis: Three new species related to Anolis nigrolineatus and A. dissimilis. Breviora (422): 1-15 - get paper here
 
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