Complicitus nigrigularis (OTA & HIKIDA, 1991)

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Higher Taxa	Agamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies
Common Names	E: Blackthroated Bloodsucker
Synonym	Calotes nigrigularis OTA & HIKIDA 1991 Bronchocela nigrigularis – MALKMUS 1994: 235 Complicitus nigrigularis — MANTHEY & GROSSMANN 1997: 166 Complicitus nigrigularis — BARTS & WILMS 2003 Complicitus nigrigularis — DAS & YAAKOB 2007
Distribution	Malaysia (Borneo, Sabah) Type locality: Kinabalu Park, Sabah, Borneo.
Reproduction	oviparous
Types	Holotype: KUZ 13223, adult male
Diagnosis	Diagnosis (genus): A relatively small agamid lizard with a snout-vent length (SVL) of < 80 mm and a tail about twice as long, which resembles Pseudocalotes species in general habitus. Easily distinguishable from these and all other agamid lizard genera by the combination of the following characters: gular sac with lateral pockets and granulated scales, crests on neck, back and along the median line of the gular pouch present, dorsals heterogeneous and tympanum visible (Manthey & Denzer 2019: 8).
Comment	Type species: Calotes nigrigularis OTA & HIKIDA 1991 is the type species of the genus Complicitus MANTHEY & GROSSMANN 1997. Abundance: only known from the type locality (Meiri et al. 2017) and from 3 specimens (Manthey & Denzer 2019).
Etymology	Named after the black coloration of the throat, from Latin niger, nigra, nigrum = black, and Latin gula = throat, also for esophagus. The genus was named after Latin complicitus = folded, referring to the lateral folds of the gular sac.
References	Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo Das, I. Lakim, M. 2008. A second specimen of Complicitus nigrigularis (Ota & Hikida, 1991). Hamadryad 32 (1): 71-73 - get paper here Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp. Malkmus,R. 1994. Herpetologische Beobachtungen aam Mt. Kinabalu, Nord-Borneo. IV. Mitt. Zool. Mus. Berlin 70: 217-249. - get paper here MANTHEY, U. & DENZER, W. 2019. Rare mountain agamas of the Kinabalu Park (Sabah, North Borneo) above 900 m a.s.l. Sauria 41 (1): 2 - 12 - get paper here Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. (ISBN 3-931587-12-6) - get paper here Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions 24 (2): 262-273 - get paper here Ota H; Hikida T 1991. Taxonomic review of the lizards of the genus Calotes Cuvier 1817 (Agamidae Squamata) from Sabah, Malaysia. Tropical Zoology 4 (2): 179-192 - get paper here
External links	IUCN Red List Google images

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