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Diploderma chapaense (BOURRET, 1937)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Japalure 
SynonymJapalura swinhonis chapaensis BOURRET 1937: 62
Japalura swinhonis chapaensis — WERMUTH 1967: 68
Japalura swinhonis chapaensis — BRYGOO 1988: 12
Japalura chapaensis — OTA 1989
Japalura chapaensis — BARTS & WILMS 2003
Japalura chapaensis — WANG et al. 2017
Diploderma chapaense — WANG et al. 2018 
DistributionN Vietnam (Chapa), China (SE and SC Yunnan)

Type locality: Chapa, Vietnam, 22° 20'N, 103° 55' E  
Reproductionoviparous 
TypesHolotype: MNHN-RA 1948.0045, female 
DiagnosisDiagnosis: Japalura chapaensis can be diagnosed from its congeners on the basis of the following set of characters: (1) body size moderate SVL 58.10–68.50 mm; (2) tail relatively long TAL/SVL 249.63% (male), 204.70%–228.61% (females); (3) limb length moderate, FLL/SVL 48.14%–54.73%, HLL/SVL 74.66%–81.41%; (4) short, w-shaped ridge present on occipital head anterior to nuchal crest; (5) tympanum concealed; (6) supraciliaries overlapping less than one-third of posterior one; (7) NSL absent (8) lateral gular fold and gular pouch present, well developed; (9) transverse gular fold absent; (10) nuchal crest scales large, crest height/HL 10.48%–14.04%; (11) MD 35–41; (12) T4S 28–30; (13) ventral scales heterogeneous in size with enlarged scales scattered ventrolaterally; (14) ground body coloration camouflaged with yellow-green, clay, and dusky brown coloration in life; (15) dorsolateral stripes present in males, jagged, sometimes discontinuous, posterolaterally away from the dorsal midline; (16) gular spot present, distinct in males, somewhat faded in females, dark spectrum yellow in both sexes; and (17) oral cavity and tongue dark spectrum yellow in life. (Wang et al. 2017)


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Comment 
EtymologyNamed after the type locality. 
References
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp.
  • Bourret,R. 1937. Notes herpétologiques sur l’indochine française. XV. Lézards et serpents reçu au laboratoire des Sciences Naturelles de l’Université au cours de l’année 1937. Descriptions de deux espèces et de deux variétés nouvelles. Bulletin Générale de l’Instruction Publique 5. Gouvernement Généneral de l’Indochine, pp. 57-82
  • Brygoo, E.R. 1988. Les types d’Agamidés (Reptiles, Sauriens) du Muséum national d’Histoire naturelle Catalogue critique. Bull. Mus. Nat. Hist. Nat. 10 (ser. 4) A (3), suppl.: 1-56
  • DENZER, WOLFGANG; ULRICH MANTHEY, PATRICK D CAMPBELL 2019. Catalogue of type specimens of the agamid lizard genus Japalura s. l. (Squamata: Agamidae: Draconinae). Zootaxa 4612 (1): 109-125 - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • MANTHEY, ULRICH; WOLFGANG DENZER, HOU MIAN & WANG XIAOHE 2012. Discovered in historical collections: Two new Japalura species (Squamata: Sauria: Agamidae) from Yulong Snow Mountains, Lijiang Prefecture, Yunnan, PR China. Zootaxa 3200: 27–48 - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Ota & WEIDENHÖFER 1992. The first male species of the poorly known agamid lizard Japalura chapensis Bourret, 1937 (Reptilia: Sauria) from northern Vietnam, with notes on its taxonomic status. Raffles Bull. Zool. 40 (2): 193-199 - get paper here
  • Ota,H. 1989. The status of an agamid lizard, Japalura swinhonis chapaensis BOURRET 1937, from Vietnam. Journal of Herpetology 23 (4): 447-450 - get paper here
  • Ota,H. 1989. A new species of Japalura (Agamidae: Lacertilia:Reptilia) from Taiwan. Copeia 1989 (3): 569-576 - get paper here
  • Poyarkov, Nikolay A.; Tan Van Nguyen, Evgeniy S. Popov,Peter Geissler, Parinya Pawangkhanant, Thy Neang, Chatmongkon Suwannapoom, Natalia B. Ananjeva, and Nikolai L. Orlov 2023. RECENT PROGRESS IN TAXONOMIC STUDIES, BIOGEOGRAPHIC ANALYSIS, AND REVISED CHECKLIST OF REPTILES IN INDOCHINA. Russian Journal of Herpetology 30 (5): 255-576 (also as vol 28 (3A): 1 – 110 [2021]) - get paper here
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Wang, Kai; Jing Che, Simin Lin, V Deepak, Datta-Roy Aniruddha, Ke Jiang, Jieqiong Jin, Hongman Chen, Cameron D Siler; 2018. Multilocus phylogeny and revised classification for mountain dragons of the genus Japalura s.l. (Reptilia: Agamidae: Draconinae) from Asia. Zoological Journal of the Linnean Society, , zly034, - get paper here
  • Wang, Kai; Ke Jiang, Yu-Fan Wang, Nikolay A. Poyarkov Jr., Jing Che, Cameron D. Siler 2017. Discovery of Japalura chapaensis Bourret, 1937 (Reptilia: Squamata: Agamidae) from Southeast Yunnan Province, China. Zoological Research 38(5): 1-11 - get paper here
 
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