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Goniurosaurus araneus GRISMER, VIETS & BOYLE, 1999

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Higher TaxaEublepharidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Vietnamese Leopard Gecko
G: Vietnamesischer Tigergecko (oder Leopardgecko)
Chinese: 越南睑虎 
SynonymGoniurosaurus araneus GRISMER, VIETS & BOYLE 1999
Goniurosaurus araneus — RÖSLER 2000: 84
Goniurosaurus araneus — KAVERKIN & SEUFER in SEUFER et al. 2005
Goniurosaurus araneus — ZHU et al. 2021 
DistributionVietnam (Cao Bang), China (Guangxi)

Type locality: 40 km SE of Cao Bang, Cao Bang Province, Vietnam.  
Reproductionoviparous 
TypesHolotype: HLMD 2572, adult male; Paratypes: ZFMK 86651 (ex-HLMD-RA-2576), ZSM 
DiagnosisDiagnosis: Goniurosaurus araneus differs from all other species of Goniurosaurus in having elongate dorsal body scales as opposed to granular body scales; and having four very wide, immaculate body bands between the nuchal loop and the caudal constriction. It differs further from G. kuroiwae and G. lichtenfelderi in having deep axillary pockets; lacking expanded and ridged occlusal tooth margins; having a greater adult SVL (115-124 mm vs. 63-97 mm), having thin, greatly elongate anterolateral and anteromedial processes of the pterygoid bone; having a posteriorly protracted nuchal loop which sits low on the nape of the neck as opposed to being rounded posteriorly and on the occiput; wide dark borders surrounding the body bands as opposed to narrow borders; and having a gracile, splayed body and limb stature as opposed to being more compact and robust. It differs further from G. kuroine in that the scales of the upper eyelids are one-half as opposed to equal in size to those on the top of the head; body tubercles do not increase in size posteriorly; claws are sheathed by four scales as opposed to being unsheathed and surrounded by 6-8 scales; having wide as opposed to narrow subdigital lamellae; having preanal pores; and the ultimate phalanges of the pes and manus are elongate and curved downward as opposed to short and straight. It differs further from G. lichtenfelderi in that it has four as opposed to three bands between the nuchal loop and the caudal constriction. It differs further from G. luii in not having a greatly enlarged row of supraorbital tubercles; higher number of preanal pores (23-29 vs. 18-22); much wider body bands and nuchal loop; nearly immaculate dorsal surfaces lacking mottling; limbs nearly immaculate, having only a few widely spaced dark blotches; dark mottling absent on lateral portions of belly; and juveniles with white as opposed to bright-orange body.bands. (Grismer et al. 1999)


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CommentCharacter table: Zhu et al. 2021

Distribution: see map in Ngo et al. 2023. 
References
  • Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp.
  • Chen, Tian-Bo; Yuan-Jun Meng, Ke Jiang,, Pi-Peng Li, Bo-Hao Wen, Wenhua Lu, James Lazell, and Mian Hou 2014. New Record of the Leopard Gecko Goniurosaurus araneus (Squamata: Eublepharidae) for China and habitat partitioning between geographically and Phylogenetically Close Leopard Geckos. IRCF Reptiles & Amphibians 21 (1): 16–27 - get paper here
  • Dickhoff, A. 2004. Ein Tiger im Terrarium - Haltung und Nachzucht von Goniurosaurus araneus (GRISMER, VIETS & BOYLE 1999). Draco 5 (18): 76-81 - get paper here
  • Grismer, L. Lee 2002. Goniurosaurus: Ancient Gekkos of the Far East. Gekko 3 (1): 22-28
  • Grismer, L. Lee, Brian E. Viets and Lawrence J. Boyle. 1999. Two new continental species of Goniurosaurus (Squamata: Eublepharidae) with a phylogeny and evolutionary classification of the genus. Journal of Herpetology 33 (3): 382-393 - get paper here
  • Grismer, L. Lee, Haitao Shi, Nicolai L. Orlov and Natalia B. Ananjeva 2002. A new species of Goniurosaurus (Squamata: Eublepharidae) from Hainan Island, China. Journal of Herpetology 36 (2): 217-224 - get paper here
  • Jonniaux P, Kumazawa Y. 2008. Molecular phylogenetic and dating analyses using mitochondrial DNA sequences of eyelid geckos (Squamata: Eublepharidae). Gene. 2008 Jan 15;407(1-2):105-15 - get paper here
  • Kaverkin, Y. 2000. Eublepharoide Geckos der Gattung Goniurosaurus BARBOUR, 1908 - Haltung und Zucht im Terrarium. Sauria 22 (4): 17-22 - get paper here
  • Ngo HN, Nguyen TQ, Phan TQ, van Schingen M, Ziegler T 2019. A case study on trade in threatened Tiger Geckos (Goniurosaurus) in Vietnam including updated information on the abundance of the Endangered G. catbaensis. Nature Conservation 33: 1–19 - get paper here
  • Ngo, H. N., Rödder, D., Grismer, L., Nguyen, T. Q., Le, M. D., Qi, S., & Ziegler, T. 2023. Extraordinary diversity among allopatric species in the genus Goniurosaurus (Squamata: Eublepharidae): understanding niche evolution and the need of conservation measures. Biodiversity and Conservation, 1-23 - get paper here
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Orlov, Nikolai L., Sergei A. Ryabov, Thien T. Nguyen, Quang T. Nguyen and Thu C. Ho. 2008. A new species of Goniurosaurus (Sauria: Gekkota: Eublepharidae) from north Vietnam. Russ. J. Herpetol. 15 (3):229-244 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Seufer, H.; Y. Kaverkin & A. Kirschner (eds.) 2005. Die Lidgeckos. Kirschner und Seufer Verlag, 238 pp.
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Wang, Ying-Yong; Jian-Huan Yang, and L. Lee Grismer 2013. A New Species of Goniurosaurus (Squamata: Eublepharidae) from Libo, Guizhou Province, China. Herpetologica 69 (2): 214-226. - get paper here
  • Wang, Ying-Yong; Jian-Huan Yang, and Rong-Feng Cui 2010. A NEW SPECIES OF GONIUROSAURUS (SQUAMATA: EUBLEPHARIDAE) FROM YINGDE, GUANGDONG PROVINCE, CHINA. Herpetologica 66 (2): 229 - get paper here
  • ZHU, X. Y., LIU, Y. J., BAI, Y., ROMÁN-PALACIOS, C. R. I. S. T. I. A. N., LI, Z., & HE, Z. Q. 2021. Goniurosaurus chengzheng sp. nov., a new species of Leopard Gecko from Guangxi, China (Squamata: Eublepharidae). Zootaxa 4996 (3): 540-554 - get paper here
  • ZHU, XIAO-YU; GUANG-YU CHEN, CRISTIAN ROMÁN-PALACIOS, ZHENG LI, ZHU-QING HE 2020. Goniurosaurus gezhi sp. nov., a new gecko species from Guangxi, China (Squamata: Eublepharidae). Zootaxa 4852 (2): 211–222 - get paper here
  • Ziegler, T.; Truong, N.Q.; Schmitz, A.; Stenke, R. & Rösler, H. 2008. A new species of Goniurosaurus from Cat Ba Island, Hai Phong, northern Vietnam (Squamata: Eublepharidae). Zootaxa 1771: 16–30 - get paper here
 
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