| Diagnosis | Diagnosis: A species of the genus Amphiesma characterized by (1) a dark dorsal background in life, dark reddish-brown or probably dark chestnut-brown in life, turning to cream, grey, pinkish-grey, tan, more or less dark greyish-brown or dark yellowish-grey in preservative, (2) on each side, a conspicuous brown or reddish-brown (cream, pale greyish-brown or pale yellowish-brown in preservative) dorsolateral stripe, narrowly edged on its both sides with dark brown or black, extending on the 5th–7th dorsal scale rows from the occipital region along the whole of the body, (3) many dorsal scales longitudinally edged with dark brown, especially on the lower part of the sides, producing a faint reticulation or irregular streaks, (4) upper edge of scales of the 1st dorsal scale row and lower edge of those of 2nd row blackish-brown, producing a very irregular ventrolateral stripe, (5) a rather narrow, irregular, blackish-brown postocular streak extending from behind the eye to the corner of the mouth, usually separated by a wide gap from the dark line edging the lower side of the dorsolateral stripe, (6) no chevron on the upper part of the neck, (7) supralabials mostly uniform, not (or barely) spotted, (8) venter uniformly cream, beige or yellowish-brown, with the outer edge of ventral tips blackish-brown, (9) 21–22 maxillary teeth, the last two moderately enlarged and separated from anterior teeth by a distinct diastema, (10) 19–19–17 dorsal scale rows, (11) dorsal scales moderately to strongly keeled on 2nd–9th dorsal and vertebral scale rows, smooth or at best feebly keeled 1st dorsal row, (12) tail moderate, proportion tail length / total length 0.221–0.252 (0.239–0.252 in males, 0.221–0.249 in females), (13) 160–173 ventrals, (14) 63–77 subcaudals, (15) anal plate divided, (16) usually 2 preoculars, and (17) 1 or 2 anterior temporals [David et al. 2015). | 
        | References | 
            Basfore, B., Kalita, M. J., Sharma, N., & Boro, A. R. 2024. An updated checklist of snakes (Reptilia: Squamata) in northeastern India derived from a review of recent literature. Journal of Threatened Taxa, 16(11), 26131-26149  - get paper hereBoulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp.  - get paper hereBoulenger, George A. 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii, 541 pp.  - get paper hereDas, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing  - get paper hereDas,I. 1996. Biogeography of the Reptiles of South Asia. Krieger Publishing Company, Malabar, Florida David P., G. Vogel & Pauwels,O. 1998. Amphiesma optatum (Hu & Djao, 1966) (Serpentes, Colubridae): an addition to the snake fauna of Vietnam, with a list of the species of the genus Amphiesma and a note on its type species. Journal of Taiwan Museum 51 (2): 83-92  - get paper hereDavid, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60  - get paper hereDavid, P.; Vogel, G. & Pauwels, O.S.G. 2005. On the occurrence of Amphiesma bitaeniatum (WALL, 1925) in Vietnam, with preliminary remarks on the group of Amphiesma parallelum (BOULENGER, 1890) (Serpentes, Colubridae, Natriciniae). Salamandra 41 (4): 167-178  - get paper hereDAVID, PATRICK; GERNOT VOGEL, TRUONG QUANG NGUYEN, NIKOLAI L. ORLOV, OLIVIER S. G. PAUWELS, ALEXANDRE TEYNIÉ, THOMAS ZIEGLER 2021. A revision of the dark-bellied, stream-dwelling snakes of the genus Hebius (Reptilia: Squamata: Natricidae) with the description of a new species from China, Vietnam and Thailand. Zootaxa 4911: 1–61  - get paper hereDAVID, PATRICK; ISHAN AGARWAL, RAMANA ATHREYA, ROSAMMA MATHEW,<br />GERNOT VOGEL & VIRAL K. MISTRY 2015. Revalidation of Natrix clerki Wall, 1925, an overlooked species in the genus Amphiesma Duméril, Bibron & Duméril, 1854 (Squamata: Natricidae). Zootaxa 3919 (2): 375–395  - get paper hereDowling, H.G., & Jenner, J.V. 1988. Snakes of Burma: checklist of reported species and bibliography. Smithsonian Herp. Inf. Serv. (76): 19 pp.  - get paper hereGUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440  - get paper hereKästle , W., Rai, K. & Schleich, H.H. 2013. FIELD GUIDE to Amphibians and Reptiles of Nepal. ARCO-Nepal e.V., 625 pp.  - get paper hereKramer, E. 1977. Zur Schlangenfauna Nepals. Revue Suisse de Zoologie 84 (3): 721-761.  - get paper hereLenz, Norbert 2012. Von Schmetterlingen und Donnerdrachen - Natur und Kultur in Bhutan. Karlsruher Naturhefte 4, Naturkundemuseum Karlsruhe, 124 pp. Mell,R. 1931. List of Chinese snakes. Lingnan Sci. Jour., Canton, 8 [1929]: 199-219. Sharma, R. C. 2004. Handbook Indian Snakes. AKHIL BOOKS, New Delhi, 292 pp. Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp. Swan, L.W., & Leviton, A.E. 1962. The herpetology of Nepal: a history, check list, and zoogeographical analysis of the herpetofauna. Proc. Cal. Acad. Sci. 32 (6) (4.s.): 103-147.  - get paper hereWall, FRANK 1925. Notes on snakes collected in Burma in 1924. J. Bombay Nat. Hist. Soc. 30 (4): 805-821.  - get paper hereWall,F. 1908. Notes on a collection of snakes from the Khasi Hills, Assam. J. Bombay nat. Hist. Soc. 18: 312-337  - get paper hereWallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Wangyal, Jigme Tshelthrim 2019. The status of herpetofauna of Bhutan. District Forest Office, District Administration, Trashigang, Bhutan, 20-39  - get paper hereZhao Ermi; Li Shengquan 1987. The herpetofaunal composition and characteristics of Mount Namjagbarwa, Tibet. Acta Herpetologica Sinica 6 (2): 36-42  - get paper hereZhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp. ZHOU, ZHENGYAN; ZHIYONG SUN, SHUO QI, YUYAN LU, ZHITONG LYU, YINGYONG WANG, PIPENG LI, JIANZHANG MA 2019. A new species of the genus Hebius (Squamata: Colubridae: Natricinae) from Hunan Province, China. Zootaxa 4674 (1): 068–082  - get paper hereZiegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225: 39-56  - get paper here |