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Podarcis raffonei (MERTENS, 1952)

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Higher TaxaLacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards)
SubspeciesPodarcis raffonei alvearioi (MERTENS 1955)
Podarcis raffonei antoninoi (MERTENS 1955)
Podarcis raffonei raffonei (MERTENS 1952) 
Common NamesE: Aeolian Wall Lizard
Italian: lucertola delle Eolie 
SynonymLacerta sicula raffonei MERTENS 1952: 313
Podarcis wagleriana raffonei — CAPULA 1987
Podarcis raffonei — CAPULA 1994
Podarcis raffonei — CORTI & LO CASCIO 2002
Podarcis raffoneae —MICHELS & BAUER 2005
Podarcis raffonei —SPEYBROECK & CROCHET 2007
Podarcis raffonei — SINDACO & JEREMČENKO 2008
Podarcis raffonei — LIVIGNI 2013: 288
Podarcis raffonei — KWET & TRAPP 2014
Podarcis raffonei — SPEYBROECK et al. 2020

Podarcis raffonei alvearioi (MERTENS 1955)
Lacerta sicula alvearioi MERTENS 1955: 31
Podarcis siculus cucchiarai PALMA 1980
Podarcis sicula alvearioi — HENLE & KLAVER 1986: 300
Lacerta sicula alvearioi — TIEDEMANN et al. 1994
Podarcis raffoneae alvearioi —MICHELS & BAUER 2005
Podarcis raffoneae cucchiarai —MICHELS & BAUER 2005
Podarcis raffonei alvearioi —LO CASCIO 2010
Podarcis raffoneae alvearioi — KWET & TRAPP 2014

Podarcis raffonei antoninoi (MERTENS 1955)
Lacerta wagleriana antoninoi MERTENS 1955: 33
Podarcis wagleriana antoninoi — BÖHME 1986: 384
Podarcis wagleriana antoninoi — ENGELMANN et al. 1993
Podarcis raffoneae antoninoi —MICHELS & BAUER 2005
Podarcis raffonei antoninoi — SINDACO & JEREMČENKO 2008
Podarcis waglerianus antoninoi — GEMEL et al. 2019 
DistributionItaly: Liparian Islands (north of Sicily), Strombolicchio Island

Type locality: ‘Eiland Strombolicchio bei Stroluboli, Liparische Inseln” (Insel Strombolischio bei Stromboli, Liparen, fide TIEDEMANN et al. 1994)

alvearioi: Italy (Liparian Islands); Type locality: Faraglione Pollara/Liparische Inseln.

antoninoi: Italy (Liparian Islands: Scoglio Faraglione); Type locality: “Halbinsel Vulcanello und auf das Tal zwischen dem Gran Cratere und dem Monte Saraceno auf Vulcano, Lipari­sche Inseln”.  
Reproductionoviparous 
TypesHolotype: SMF 43919, adult male; collected by Antonino TRISCHITTA, 31 May 1951.
Holotype: SMF 48447; paratypes NMW 15745:1-4 [antoninoi]
Holotype: SMF 48296; paraypes: NMW 15746:1,2 [alvearioi]
Holotype: MZUP (Museo Zoologico dell’Università di Palermo), male; paratypes: BMNH, MZUF [cucchiarai] 
DiagnosisDiagnosis: “Eine qroße, stark verdüsterte Inselrasse von Lacerta sicula, ohne jede Neigung zur Blaufärbung. Oberseite dunkelbraun mit undeutlicher sepiafarbener Fleckenzeichnung, Unterseite rauchgrau rnit türkisblauen Fleckchen auf der äußerstcn Ventraliareihe; kein blauer Axillarfleck. Pileus braun mit zahlreichen schwarzen Flecken. Massetericum fehlt. Länge von Kopf + Rumpf : 85 mm, Länge des Schwanzes gelegentlich die doppeite Kopf-Rumpf-Länge überschreitend.” (Mertens 1952)


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CommentGenome: Gabrielli et al. 2023

Synonymy: The validity of the sub-species raffonei (Mertens 1952), alvearioi (Mertens 1955), cucchiarai Di Palma 1980 and antoninoi (Mertens 1955) is not supported by genetic data (Capula, 2004). 
Etymology"Benannt ist die Rasse, dem Wunsche des Sammlers entsprechend, nach RAFFONE, dem Familiennamen seiner verstorbenen Frau." [= The race has been named, according to the wish of his collector, after RAFFONE, the maiden name of his late wife]. The name therefore has to be P. raffoneae according to MICHELS & BAUER 2005. However, Oscar Arribas (pers. comm., 26 Jul 2016) objected that “The Code [Article 32.5.1] states that gender and species must concordate in grammatical gender, but none about correcting if the sex of the hommaged person is not correctly declined. Raffonae is [thus] an unjustified enmendation.” 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Böhme,W. 1986. Podarcis wagleriana - Sizilianische Mauereidechse. In: Böhme,W. (ed.), Handbuch der Reptilien und Amphibien Europas, Band 2/II., Echsen III (Podarcis). Aula-Verlag Wiesbaden, pp. 377-387
  • Capula, M. 2006. Population heterogeneity and conservation of the Aeolian wall lizard, Podarcis raffonei. In: Mainland and insular lacertid lizards: a mediterranean perspective, Corti,C., Lo Cascio, P., Biaggini, M., Eds. Firenze University Press, Firenze, p. 24-32
  • Capula, M., Luiselli, L., Bologna, M.A., Ceccarelli, A. 2002. The decline of the Aeolian wall lizard, Podarcis raffonei: causes and conservation proposals. Oryx 36: 66-72 - get paper here
  • Capula, Massimo, Claudia Corti, Pietro Lo Cascio and Luca Luiselli. 2014. Thermal ecology of the Aeolian Wall Lizard, Podarcis raffonei. What about body temperatures in microinsular lizards? Pp. 39-47. In: M. Capula and C. Corti (eds.) Scripta Herpetologica: Studies on Amphibians and reptiles in honour of Benedetto Lanza. Edizioni Belvedere, Latina
  • Capula,M. 1994. Allozyme variation and differentiation in the lacertid lizard, Podarcis wagleriana (Reptilia: Lacertidae). Biol. J. Linn. Soc. 52: 177-196 - get paper here
  • Cascio, Pietro Lo 2010. ATTUALI CONOSCENZE E MISURE DI CONSERVAZIONE PER LE POPOLAZIONI RELITTE DELL’ENDEMICA LUCERTOLA DELLE EOLIE, PODARCIS RAFFONEI (Squamata Sauria). Naturalista sicil., S. IV, XXXIV (3-4): 295-317 - get paper here
  • Corti, C. & Lo Cascio, P. 2002. The lizards of Italy and Adjacent Areas. Edition Chimaira, Frankfurt, 165 pp.
  • D’Amico, Marcello; Giulia Bastianelli, Francesco Paolo Faraone, and Mario Lo Valvo 2018. The Spreading of the Invasive Italian Wall Lizard on Vulcano, the Last Island Inhabited by the Critically Endangered Aeolian Wall Lizard. Herp. Cons. Biol. 13 (1): 146–157 - get paper here
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • Gabrielli, M., Benazzo, A., Biello, R., Ancona, L., Fuselli, S., Iannucci, A., ... & Bertorelle, G. 2023. A high-quality reference genome for the critically endangered Aeolian wall lizard, Podarcis raffonei. Journal of Heredity, esad014 - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248 - get paper here
  • Gippoliti, Spartaco; Massimo Capula, Gentile Ficetola, Daniele Salvi, and Franco Andreone 2017. Threatened by Legislative Conservationism? The Case of the Critically Endangered Aeolian Lizard. Front. Ecol. Evol., 20 - get paper here
  • Kwet, A. 2010. Reptilien und Amphibien Europas. Kosmos-Verlag, Stuttgart, 252 pp.
  • Kwet, Axel & Benny Trapp 2014. Reptilien in Europa – eine aktuelle Übersicht. Draco 15 (60): 6-25 - get paper here
  • Kwet, Axel & Benny Trapp 2014. Liste der Reptilien Europas. Draco 15 (60): 72-79 - get paper here
  • LiVigni, F. (ed.) 2013. A Life for Reptiles and Amphibians, Volume 1. Chimaira, Frankfurt, 495 pp. - get paper here
  • Lo Cascio, Pietro & Massimo Capula 2011. Does diet in lacertid lizards reflect prey availability? Evidence for selective predation in the Aeolian wall lizard, Podarcis raffonei (Mertens, 1952) (Reptilia, Lacertidae). Biodiversity Journal 2 (2): 89-96 - get paper here
  • LO CASCIO, PIETRO; FLAVIA GRITA, LIVIA GUARINO & CLAUDIA SPECIALE 2014. A LITTLE IS BETTER THAN NONE: NEW INSIGHTS INTO THE NATURAL HISTORY OF THE AEOLIAN WALL LIZARD PODARCIS RAFFONEI FROM LA CANNA STACK (Squamata Sauria). Naturalista sicil., S. IV, XXXVIII (2): 203-214
  • Mertens, R. 1952. Neue Eidechsenrassen von den Liparischen Inseln. Senckenbergiana 32: 309-314
  • Mertens, R. 1955. Die Mauereidechsen der Liparischen Inseln, gesammelt von Dr. Antonino Trischitta. Senckenbergiana Biologica 36: 25-40
  • Michels, J.P. & A.M. Bauer 2004. Some corrections to the scientific names of amphibians and reptiles. Bonner Zoologische Beiträge 52: 83–94 [2005] [Note: Dubois 2007 rejected most of the changes made in this paper] - get paper here
  • Palma, Maria Gabriella di 1980. La Lucertola del Faraglione “La Canna” nelle Isole Eolie: Podarcis sicula cucchiarai subsp. nova (Reptilia, Lacertidae). Il Naturalista Siciliano (IV) IV (1-2): 3-12 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Speybroeck, J. & Crochet, P.A. 2007. Species list of the European herpetofauna - a tentative update. Podarcis 8 (1/2): 8-34 - get paper here
  • Tiedemann,F.; Häupl,M. & Grillitsch,H. 1994. Katalog der Typen der herpetologischen Sammlung nach dem Stand vom 1. Jänner 1994. Teil II: Reptilia. Kat. wiss. Samml. Naturhist. Mus. Wien 10 (Vertebrata 4): 1-110
 
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