Smaug breyeri (VAN DAM, 1921)

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Higher Taxa	Cordylidae (Cordylinae), Scincoidea, Sauria, Squamata (lizards)
Subspecies
Common Names	E: Waterberg Dragon Lizard, Waterberg Girdled Lizard
Synonym	Zonurus breyeri VAN DAM 1921: 239 Cordylus warreni breyeri — FITZSIMONS 1943 Cordylus breyeri — BRANCH 1998 Smaug breyeri — STANLEY et al. 2011 Smaug breyeri — BATES et al. 2014: 208
Distribution	South Africa: Northern Province - Waterberg and environs Type locality: Farm Geelhoutkop, Waterberg District [Northern Province, South Africa].
Reproduction	Ovoviviparous.
Types	Holotype: TM 3769, adult female, paratypes: TM
Diagnosis	Unfortunately we had to temporarily remove additional information as this was scraped by multiple AI companies who sell that data to their customers. These details, e.g. detailed descriptions or comparisons (about 2908 characters), are available for collaborators and contributors. Please contact us for details.
Comment	See also S. warreni.
Etymology	Named after Dr. Hermann Gottfried Breyer (1864-1923), a Dutch naturalist, botanist, and physician who collected in Transvaal and Mozambique (1890-1910). He became Curator (1901) and then Director (1913-1921) of the renamed Transvaal Museum and Zoological Gardens.
References	Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp. Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Branch, W. R. 1998. Field Guide to the Snakes and Other Reptiles of Southern Africa. 3rd ed. Fully Revised and Updated to Include 83 New Species. Ralph Curtis Books (Sanibel Island, Florida), 399 pp. Broeckhoven, C., Diedericks, G., Hui, C., Makhubo, B. G. and Mouton, P. l. F. N. 2016. Enemy at the gates: Rapid defensive trait diversification in an adaptive radiation of lizards. Evolution, 70: 2647–2656.doi:10.1111/evo.13062 - get paper here Dam, G.P.F. van 1921. Descriptions of new species of Zonurus, and notes on the species of Zonurus occurring in the Transvaal. Annals Transvaal Mus. 7 (4): 239-243 - get paper here FitzSimons, V.F. 1943. The lizards of South Africa. Transvaal Museum Memoir No.1 (Pretoria), 528 pp. Mouton, P le Fras N; Alexander Flemming; Michael Bates and Chris Broeckhoven 2018. The relationship between generation gland morphology and armour in Dragon Lizards (Smaug): a reassessment of ancestral states for the Cordylidae. Amphibia-Reptilia 39 (4): 445–456 - get paper here Reissig, J. 2014. Girdled Lizards and Their Relatives. Edition Chimaira, Frankfurt, ISBN-10: 3899734378 Stanley, E. L. and Bates, M. F. 2014. Here be dragons: a phylogenetic and biogeographical study of the Smaug warreni species complex (Squamata: Cordylidae) in southern Africa. Zoological Journal of the Linnean Society 172 (4): 892–909; doi: 10.1111/zoj.12187 - get paper here Stanley, Edward L.; Aaron M. Bauer; Todd R. Jackman, William R. Branch, P. Le Fras N. Mouton 2011. Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): 53-70. - get paper here Tolley, K.A., Conradie, W., Pietersen, D.W., Weeber, J., Burger, M. & Alexander, G.J. (eds) 2023. Conservation status of the reptiles of South Africa, Eswatini and Lesotho. Suricata 10. South African National Biodiversity Institute, Pretoria - get paper here
External links	IUCN Red List National Center for Biotechnology Information Google images

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