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Tricheilostoma broadleyi (WALLACH & HAHN, 1997)

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Higher TaxaLeptotyphlopidae, Epictinae, Epictini, Typhlopoidea, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymLeptotyphlops broadleyi WALLACH & HAHN 1997
Leptotyphlops broadleyi — MCDIARMID, CAMPBELL & TOURÉ 1999: 24
Guinea broadleyi — ADALSTEINSSON, BRANCH, TRAPE, VITT & HEDGES 2009
Tricheilostoma broadleyi —HEDGES 2011
Tricheilostoma broadleyi — WALLACH et al. 2014: 737 
DistributionSC Ivory Coast (Cote d'Ivoire), 300 - 500 m elevation

Type locality: Tropical Ecology Station, Lamto, 300 m north of the Bandama River, Toumodi Department, Cote d'Ivoire.  
Reproductionoviparous 
TypesHolotype: MNHN-RA 1977.0103; paratypes: MNHN-RA 1977.0092-0102 
DiagnosisDiagnosis: A diminutive species of Leptotyphlops with a maximum length less than 115 mm that is closely related to L. bicolor but differs from it in the following characteristics: lower number of middorsal scales (17 - 19, x = 186. vs. 247 - 288, x = 262.), higher number of subcaudals (1 - 16, x = 1.9 vs. 8 - 1, x = 10.2), size and shape of prefrontal (broader than deep, > 0.5 head width vs. as deep as broad, < 0.5 head width), tail termination (vertically compressed ridge vs. cone), lower length/width ratio (24 - 44, x = 5.6 vs. 8 - 69, x = 52.9), longer tail/total length ratio (5 .1 - 7.0, x = 5.6 vs. 2.4 - 4.0, x = .2), and smaller maximum size (115 vs. 185 mm).


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Comment 
EtymologyNamed in honor of Donald G. Broadley (1932-2016), famous African herpetologist. Broadley died on the same day as Benedetto Lanza (10 March 2016). See Bates & Branch 2018 for an obituary. 
References
  • Adalsteinsson, S.A.; Branch, W.R.; Trapé, S.; Vitt, L.J. & Hedges, S.B. 2009. Molecular phylogeny, classification, and biogeography of snakes of the Family Leptotyphlopidae (Reptilia, Squamata). Zootaxa 2244: 1-50 - get paper here
  • Bates, Michael F. & William R. Branch 2018. End of an era for African Herpetology—An obituary for a remarkable man: Donald George Broadley (29 May 1932 – 10 March 2016). African Journal of Herpetology, DOI: 10.1080/21564574.2018.1437085 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Chippaux, Jean-Philippe & Kate Jackson 2019. Snakes of Central and Western Africa. Johns Hopkins University Press, 448 pp. [detaileld review in HR 51 (1): 161] - get paper here
  • Hahn D. E. & V. WALLACH, 1998. Comments on the systematics of Old World Leptotyphlops (Serpentes: Leptotyphlopidae), with description of a new species. Hamadryad 23: 50-62 - get paper here
  • HEDGES, S. BLAIR 2011. The type species of the threadsnake genus Tricheilostoma Jan revisited (Squamata, Leptotyphlopidae). Zootaxa 3027: 63–64 - get paper here
  • McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp.
  • Pauwels, O.S.G.; Brecko, J. 2023. Book review: Guide des Serpents d'Afrique Occidentale, Centrale et d'Afrique du Nord. Jean-François Trape.IRD Editions, Marseille, France (www.editions.ird.fr). 896 pp. Herpetological Review 54 (4): 700-702.
  • Trape, J.-F. 2023. Guide des serpents d’Afrique occidentale, centrale et d’Afrique du Nord. IRD Éditions, Marseille, 896 pp.
  • TRAPE, JEAN-FRANÇOIS & CELLOU BALDÉ 2014. A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa 3900 (3): 301–338 - get paper here
  • Wallach, V. & D. E. Hahn 1997. Leptotyphlops broadleyi, a new species of worm snake from Cote d'Ivoire (Serpentes: Leptotyphlopidae). African Journal of Herpetology 46 (2): 103-109. - get paper here
  • Wallach, Van; Kenneth L. Williams, Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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