Bothrocophias lojanus (PARKER, 1930)

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Higher Taxa	Viperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names	E: Lojan Lancehead
Synonym	Bothrops lojanus PARKER 1930 Bothrops lojana PARKER 1930 Bothrops alticola PARKER 1939 Bothriopsis alticola — WELCH 1994: 31 Bothrops lojanus — WELCH 1994: 33 Bothrops lojanus — MCDIARMID, CAMPBELL & TOURÉ 1999: 266 Bothrops lojanus — CARRASCO et al. 2012 Bothrops lojanus — WALLACH et al. 2014: 117 Bothrocophias lojanus — HAMDAN et al. 2019
Distribution	Ecuador (Loja, Zamora-Chinchipe), Peru; elevation up to 2500 m or higher. Type locality: Loja, Provincia Loja, Ecuador, 2200 m elevation.
Reproduction	ovovivparous
Types	Holotype: BMNH 1930.1.30.1
Diagnosis	Additional details, e.g. a detailed description or comparisons (2133 characters), are available for collaborators and contributors. Please contact us for details.
Comment	Venomous! Incertae sedis fide FENWICK et al. 2009. Bothrops lojanus nests within Bothrocophias (Alencar et al. 2016) although support for a position within the latter is not very strong. Conservation: one of the 30 most endangered viper species (Maritz et al. 2016).
Etymology	Named after the type locality.
References	Alencar, Laura R.V.; Tiago B. Quental, Felipe G. Grazziotin, Michael L. Alfaro, Marcio Martins, Mericien Venzon, Hussam Zaher 2016. Diversification in vipers: Phylogenetic relationships, time of divergence and shifts in speciation rates. Molecular Phylogenetics and Evolution 105: 50–62 - get paper here Campbell, J.A. & Lamar, W.W. 1989. The Venomous Reptiles of Latin America. Comstock Publishing/Cornell University Press, Ithaca Carrasco, P.A., C.I. Mattoni, G.C. Leynaud, and G.J. Scrocchi. 2012. Morphology, phylogeny and taxonomy of South American bothropoid pitvipers (Serpentes, Viperidae). Zoologica Scripta 41: 109-124 - get paper here Carrasco, Paola A; Pablo J Venegas, Valencia, Jorge H 2016. First confirmed records of the endangered Andean pitviper Bothrops lojanus Parker, 1930 (Viperidae: Crotalinae) from Peru. Herpetology Notes 9: 297-301 - get paper here FENWICK, ALLYSON M.; RONALD L. GUTBERLET JR, JENNAFER A. EVANS, CHRISTOPHER L. PARKINSON 2009. Morphological and molecular evidence for phylogeny and classification of South American pitvipers, genera Bothrops, Bothriopsis, and Bothrocophias (Serpentes: Viperidae). Zoological Journal of the Linnean Society 156 (3): 617-640 - get paper here Hamdan, B, Guedes, TB, Carrasco, PA, Melville, J. 2019. A complex biogeographic history of diversification in Neotropical lancehead pitvipers (Serpentes, Viperidae). Zoologica Scripta 49: 145– 158 [print 2020] - get paper here Maritz, Bryan; Johannes Penner, Marcio Martins, Jelka Crnobrnja-Isailović, Stephen Spear, Laura R.V. Alencar, Jesús Sigala-Rodriguez, Kevin Messenger, Rulon W. Clark, Pritpal Soorae, Luca Luiselli, Chris Jenkins, Harry W. Greene 2016. Identifying global priorities for the conservation of vipers. Biological Conservation, doi:10.1016/j.biocon.2016.05.004 - get paper here McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp. OLIVEIRA-DALLAND, LUIS G.; LAURA R.V. ALENCAR, LEANDRO R. TAMBOSI, PAOLA A. CARRASCO, RHETT M. RAUTSAW, JESUS SIGALA-RODRIGUEZ, GUSTAVO SCROCCHI & MARCIO MARTINS. 2022. Conservation gaps for Neotropical vipers: Mismatches between protected areas, species richness and evolutionary distinctiveness. Biological Conservation 275(109750). - get paper here Parker,H.W. 1930. Two new reptiles from southern Ecuador. Ann. Mag. nat. Hist. (10) 5: 568-571 - get paper here Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here Valencia, J. H., K. Garzón-Tello & M. E. Barragán-Paladines 2016. Serpientes venenosas del Ecuador: sistemática, taxonomía, historia natural, conservación, envenenamiento y aspectos antropológicos. Quito, Ecuador, Fundación Herpetológica Gustavo Orcés, Universidad de Texas, Fondo Ambiental Nacional, 652 pp. [review in HR 49 (1): 152, 2018] Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
External links	IUCN Red List National Center for Biotechnology Information Google images

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