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Cordylus machadoi LAURENT, 1964

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Higher TaxaCordylidae (Cordylinae), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Machado’s Girdled Lizard, Kaokoveld Girdled Lizard
G: Machado’s Gürtelschweif 
SynonymCordylus vittifer machadoi LAURENT 1964: 49
Cordylus machadoi — BRANCH 1998
Cordylus machadoi — STANLEY et al. 2011
Cordylus machadoi — REISSIG 2014
Cordylus machadoi — SCHLEICHER 2020 
DistributionS Angola, N Namibia

Type locality: Leba, Humpata, environs de Sá da Bandeira, elevation 1800 m, Huila.  
ReproductionOvoviviparous. 
TypesHolotype: MD = Museu Dundo; paratype: MCZ 
DiagnosisDiagnosis: “Race de Cordylus vittifer différant de la forme typique par son plus grand nombre de rangées longitudinales d'écailles dorsales: 26-28 au lieu de 20-24 et la rostrale moins large (hauteur comprise 2,34 à 2,32 fois dans sa largeur au lieu de 3 à 4 fois)” (Laurent 1964).


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CommentDistribution: see maps in GREENBAUM et al. 2012, Lobon-Rovira et al. 2025: 14 (Fig. 10)

Cordylus v. machadoi was originally distinguished from other C. vittifer on the basis of its greater numbers of longitudinal dorsal scale rows (26–28 versus 20–24) and a narrower rostral scale (2.32–2.34 times wider than deep versus 3–4 times wider). Reichenow (1887), in his original description of Zonurus vittifer (Fig. S2), reported only 20–22 longitudinal dorsal scale rows and did not mention the size or shape of the rostral, and it therefore seems that Laurent (1964) was referring to Loveridge (1944a, see table on p. 15 in Bates et al. 2023) and possibly also FitzSimons (1943) when comparing machadoi to vittifer. 
EtymologyNamed after Dr. A. de Barros-Machado, who collected the type in Angola. 
References
  • Baptista NL, António T, Branch WR. 2019. The herpetofauna of Bicuar National Park and surroundings, southwestern Angola: a preliminary checklist. Amphibian & Reptile Conservation 13(2) [Special Section]: 96–130 (e203) - get paper here
  • Bates MF, Lobón-Rovira J, Stanley EL, Branch WR, Vaz Pinto P 2023. A new species of green-eyed Cordylus Laurenti, 1768 from the west-central highlands of Angola, and the rediscovery of Cordylus angolensis (Bocage, 1895) (Squamata: Cordylidae). Vertebrate Zoology 73: 599-646 - get paper here
  • Branch, W. R. 1998. Field Guide to the Snakes and Other Reptiles of Southern Africa. 3rd ed. Fully Revised and Updated to Include 83 New Species. Ralph Curtis Books (Sanibel Island, Florida), 399 pp.
  • Butler, B O; Ceríaco, L M P; Marques, M P; Bandeira, S; Júlio, T; Heinicke, M P & Bauer, A M; 2019. Herpetological survey of Huíla Province, Southwest Angola, including first records of Bicuar National Park. Herpetological Review 50 (2): 225-240 - get paper here
  • Ceríaco, Luis M. P.; Mariana P. Marques, Ilunga André, Esteves Afonso, David C. Blackburn & Aaron M. Bauer 2020. ILLUSTRATED TYPE CATALOGUE OF THE “LOST” HERPETOLOGICAL COLLECTIONS OF MUSEU DO DUNDO, ANGOLA. Bulletin of the Museum of Comparative Zoology 162 (7): 379-440 - get paper here
  • Conradie, Werner; Brain Reeves, Sandile Mdoko, Lwandiso Pamla, Oyama Gxabhu 2020. Herpetological survey of the Ongeluksnek (Malekgalonyane) Nature Reserve on the foothills of the Drakensberg, Eastern Cape Province, South Africa. Herpetology Notes 13: 717-730 - get paper here
  • Greenbaum, Eli; Edward L. Stanley, Chifundera Kusamba, Wandege M. Moninga, Stephen R. Goldberg & Charles R. Bursey 2012. A new species of Cordylus (Squamata: Cordylidae) from the Marungu Plateau of south-eastern Democratic Republic of the Congo. African Journal of Herpetology 61 (1): 14-39 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • Laurent, R.F. 1964. Reptiles et batraciens de l'Angola (troisième contribution). Companhia de Di amantes de Angola (Diamang), Serviços Culturais, Museu do Dundo (Angola), No. 67, 165 pp.
  • Lobón-Rovira, J., Vaz Pinto, P., Keates, C., & Stanley, E. L. 2025. Ancient polymorphism, secondary contact, taxonomic inflation or all of them together? Mito-nuclear discordance in the Angolan girdled lizard (Cordylidae: Cordylus), with the description of a new species from the central highlands. Zoological Journal of the Linnean Society, 205(1), zlaf112 - get paper here
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • MARQUES, MARIANA P.; LUIS M. P. CERÍACO, EDWARD L. STANLEY, SUZANA A. BANDEIRA, ISHAN AGARWAL, AARON M. BAUER 2019. A new species of Girdled Lizard (Squamata: Cordylidae) from the Serra da Neve Inselberg, Namibe Province, southwestern Angola. Zootaxa 4668 (4): 503–524 - get paper here
  • Reissig, J. 2014. Girdled Lizards and Their Relatives. Edition Chimaira, Frankfurt, ISBN-10: 3899734378
  • Schleicher, Alfred 2020. Reptiles of Namibia. Kuiseb Publishers, Windhoek, Namibia, 271 pp.
  • Stanley, Edward L.; Aaron M. Bauer; Todd R. Jackman, William R. Branch, P. Le Fras N. Mouton 2011. Between a rock and a hard polytomy: rapid radiation in the rupicolous girdled lizard (Squamata: Cordylidae). Molecular Phylogenetics and Evolution, 58(1): 53-70. - get paper here
  • STANLEY, EDWARD L.; LUIS M. P. CERÍACO, SUZANA BANDEIRA, HILARIA VALERIO, MICHAEL F. BATES & WILLIAM R. BRANCH 2016. A review of Cordylus machadoi (Squamata: Cordylidae) in southwestern Angola, with the description of a new species from the Pro-Namib desert,. Zootaxa 4061 (3): 201–226 - get paper here
 
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