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Acanthodactylus hardyi HAAS, 1957

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
Subspecies 
Common NamesE: Hardy’s Fringe-fingered Lizard 
SynonymAcanthodactylus scutellatus hardyi HAAS, 1957: 72
Acanthodactylus scutellatus hardyi — BONS & GIROT 1964
Acanthodactylus scutellatus hardyi — SALVADOR 1982: 119
Acanthodactylus hardyi — HARRIS & ARNOLD 2000
Acanthodactylus scutellatus hardyi — CROCHET et al. 2003
Acanthodactylus hardyi — RIFAI et al. 2003
Acanthodactylus hardyi — SCHLÜTER 2010: 62
Acanthodactylus scutellatus hardyi — MOHAMMED et al. 2017 
DistributionJordan, Iraq, Saudi Arabia, Kuwait.

Type locality: “Hirmas Station, Saudi Arabia”.  
Reproductionoviparous. 
TypesHolotype: HUJ Hebrew University of Jerusalem 2682 
DiagnosisDiagnosis: Smaller than typical subspecies. Relatively wider head with nasal regions more raised. Larger gulars. Fragmented fourth supraocular. 12, sometimes 14, almost straight longitudinal rows of ventrals. Dorsals flat, granular, and never keeled. Temporals large and flat (Salvador 1982: 121). 
Comment 
EtymologyNamed after Dr. J. E. Hardy, head of the Departrnent of Entomology, British Mandatory Government, Palestine (1942) and collector of the holotype. 
References
  • Aloufi, A., Al Zayer, M., & Amr, Z. S. 2023. Reptiles and Amphibians along the Coastal Area of the Eastern Province, Saudi Arabia. Ecologica Montenegrina, 68, 66-87 - get paper here
  • Baha El Din, Sherif M. 2007. A new lizard of the Acanthodactylus scutellatus group (Squamata: Lacertidae) from Egypt. Zoology in the Middle East 40: 21-32 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Blanc CP, Ineich I. 1985. Etude sur les acanthodactyles de Tunisie: VII – Les acanthodactyles de l’extrême-sud tunisien. Amphibia-Reptilia 6: 45–52 - get paper here
  • Bons,J. & GIROT,B. 1964. Révision de l'espèce Acanthodactylus scutellatus (Lacertidé-Saurien). Bull. Soc. Sci. nat. phys. Maroc 42: 311-334 [1962]
  • Boulenger, G.A. 1878. Sur les espèces d'Acanthodactylus des bords de la Mediterranée. Bull. Soc. zool. France 3: 179-197 - get paper here
  • Boulenger, G.A. 1919. Sur les lézards du genre Acanthodactylus Wieg. Bull. Soc. zool. France 43: 143-155 [1918] - get paper here
  • Boulenger, George A. 1891. Catalogue of the reptiles and batrachians of Barbary (Morocco, Algeria, Tunisia), based chiefly upon the notes and collections made in 1880-1884 by M. Fernand Lataste. Tr. Zool. Soc. 13: 93-164
  • Cooper Jr., W.E. 2005. Duration of movement as a lizard foraging movement variable. Herpetologica 61 (4): 363-372 - get paper here
  • Crochet, P.A.; Geniez, P. & Ineich, I. 2003. A multivariate analysis of the fringe-toed lizards of the Acanthodactylus scutellatus group (Squamata: Lacertidae): systematic and biogeographical implications. Zool. J. Linnean Soc.137: 117–155 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1839. Erpétologie Générale on Histoire Naturelle Complète des Reptiles. Vol. 5. Roret/Fain et Thunot, Paris, 871 pp. - get paper here
  • GRAMENTZ, D. 2002. Zur Ökologie und zum Verhalten von Acanthodactylus scutellatus (AUDOUIN, 1809) in West-Tunesien. Die Eidechse 13 (1): 14-21 - get paper here
  • Haas G. and Werner, Y. L. 1969. Lizards and snakes from Southwestern Asia, collected by Henry Field. Bull. Mus. Comp. Zool. Harvard 138: 327-406. - get paper here
  • Haas, Georg 1957. Some amphibians and reptiles from Arabia. Proc. Cal. Acad. Sci. 29 (3): 47-86 - get paper here
  • Johann, H. 1981. Herpetologische Eindrücke auf einer Reise durch die Sahara. Herpetofauna 3 (13): 17-21 - get paper here
  • KRONIGER, M. 1995. Zur Haltung von Acanthodactylus scutellatus (AUDOUIN, 1809). Die Eidechse 6 (14): 1-3 - get paper here
  • Loveridge, A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Mellado,J. & Olmedo,G. 1990. El género Acanthodactylus en Marruecos: problemas de identificación en los grupos de especies A. pardalis y A. scutellatus. Amphibia-Reptilia 11: 131-146 - get paper here
  • Padial, J. M. 2006. COMMENTED DISTRIBUTIONAL LIST OF THE REPTILES OF MAURITANIA (WEST AFRICA). Graellsia, 62(2): 159-178 - get paper here
  • Rifai, L., D. Modrý, P. Nečas & Z. S. Amr 2003. The occurrence of Acanthodactylus hardyi Haas, 1957 in the Hashemite Kingdom of Jordan and notes on its ecology. Zoology in the Middle East 28: 33 - get paper here
  • Salvador, A 1982. A revision of the lizards of the genus Acanthodactylus (Sauria: Lacertidae). Bonner Zool. Monogr. (16): 1-167 - get paper here
  • Schleich, H.H., Kästle,W., Kabisch, K. 1996. Amphibians and Reptiles of North Africa. Koeltz, Koenigstein, 627 pp.
  • Schlüter, U. 2007. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. Teil 2: Gundlagen der Terrarienhaltung und Reproduktionsbiologie. Reptilia (Münster) 12 (67): 56-65 - get paper here
  • Schlüter, U. 2010. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. Teil 5b: Die Acanthodactylus-scutellatus-Gruppe. Reptilia (Münster) 15 (84): 62-66 - get paper here
  • Schlüter, U. 2010. Fransenfingereidechsen (Acanthodactylus) in der Natur und im Terrarium. Teil 5a: Die Acanthodactylus-scutellatus-Gruppe. Reptilia (Münster) 15 (83): 40-45 - get paper here
  • Schmidt, K. P. & Marx, H. 1956. The herpetology of Sinai. Fieldiana 39 (4): 21-40 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • van der Kuyl, Antoinette C.; Donato L. Ph. Ballasina, John T. Dekker, Jolanda Maas, Ronald E. Willemsen and Jaap Goudsmit 2002. Phylogenetic Relationships among the Species of the Genus Testudo (Testudines: Testudinidae) Inferred from Mitochondrial 12S rRNA Gene Sequences. Molecular Phylogenetics and Evolution 22: 174-183 - get paper here
  • Zulueta, A. de 1909. Nota sobre reptiles de Cabo Juby (N.W. de Africa). Bol. Real Sociedad Esp. Hist. Nat. 9: 354-355
 
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