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Acanthosaura titiwangsaensis WOOD, GRISMER, GRISMER, AHMAD, ONN & BAUER, 2009

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Malayan Mountain Horned Agamid, Titiwangsa Horned Tree Lizard 
SynonymAcanthosaura titiwangsaensis WOOD, GRISMER, GRISMER, AHMAD, ONN & BAUER 2009
Acanthosaura titiwangsaensis — GRISMER & QUAH 2019 
DistributionW Malaysia (Pahang)

Type locality: Bishop’s Trail, Fraser’s Hill, Pahang, Peninsular Malaysia (3º43.230'N; 101º44.887'E), 1,112 m elevation.  
Reproduction 
TypesHolotype: ZRC 2.6805, adult male, collected on 28 June 2008 by Jesse L. Grismer, L. Lee Grismer, Perry L. Wood Jr., at 2330 h.; Paratypes: ZFMK and ZRC 
DiagnosisDiagnosis: Maximum SVL of adult males 118.4 mm, maximum SVL of females 112.7 mm; head triangular, snout short (SL/HL 46–52%); eye moderate (EYE/HL 28–30%); no enlarged elliptical scales below orbit; scales on the canthus rostralis-supraorbital ridge 14 or 15; large rectangular rostral scale; a short cylindrical spine above posterior margin of eye surrounded by a rosette of four smaller keeled scales at base; spine on occiput between tympanum and nuchal crest present; tympanum naked, oblong, taller than wide; dewlap present in males and females; body triangular in cross section; scales of flanks intermixed with randomly arranged clusters of larger keeled scales; nuchal crest present; 11–13 scales in nape diastema; vertebral crest composed of enlarged, pointed scales beginning at shoulder region and tapering to a series of enlarged keeled scales on anterior portion of tail; crest scales largest at base of nape; tail 140–150% of SVL; black collar present; posterior 2/3 of tail black to dark brown; greenish-yellow to light orange spots encircled by gray on body, gular region orangish yellow; no yellow medial stripe on dewlap.


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Comment 
EtymologyThis specific epithet titwangsaensis is in reference to the mountain range, Banjaran Titiwangsa, to which we believe this species is likely to be restricted. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Liu S, Rao D 2019. A new species of the genus Acanthosaura from Yunnan, China (Squamata, Agamidae). ZooKeys 888: 105-132 - get paper here
  • NORHAYATI, A.; FARAH, A.D.; CHAN, K.O.; DAICUS, B. and MUIN, M.A. 2011. AN UPDATE OF HERPETOFAUNAL RECORDS FROM BUKIT FRASER, PAHANG, PENINSULAR MALAYSIA. Malays. Appl. Biol. (2011) 40(2): 9–17 - get paper here
  • PAUWELS, OLIVIER S. G.; MONTRI SUMONTHA, KIRATI KUNYA, AWAT NITIKUL, PHAMON SAMPHANTHAMIT, PERRY L. WOOD, JR. & LEE L. GRISMER 2015. Acanthosaura phuketensis (Squamata: Agamidae), a new long-horned tree agamid from southwestern Thailand. Zootaxa 4020 (3): 473–494 - get paper here
  • WOOD JR., PERRY L.;JESSE L. GRISMER, L. LEE GRISMER, NORHAYATI AHMAD, & CHAN KIN ONN & AARON M. BAUER 2009. Two new montane species of Acanthosaura Gray, 1831 (Squamata: Agamidae) from Peninsular Malaysia. Zootaxa 2012: 28-46 - get paper here
  • Wood, P.L.; Grismer, L.L.; Grismer, J.L.; Neang, T.; Chav, T. & holden, J. 2010. A new cryptic species of Acanthosaura Gray, 1831 (Squamata: Agamidae) from Thailand and Cambodia. Zootaxa 2488: 22–38 - get paper here
 
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