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Afroedura bogerti LOVERIDGE, 1944

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Namba Flat Gecko, Bogert's Rock Gecko 
SynonymAfroedura karroica bogerti LOVERIDGE 1944: 1
Afroedura bogerti — ONDERSTALL 1984
Afroedura bogerti — RÖSLER 2000: 57
Afroedura bogerti — BAUER 2013 (plates)
Afroedura cf. bogerti — HERRMANN & BRANCH 2013
Afroedura cf. bogerti — BUTLER et al. 2019 
DistributionEC Angola (Cuanza), Namibia

Type locality: Namba (Mombolo), Cuanza Sul Province, Angola [Mombolo: 12°10’S, 14°50’E, Missão da Namba: 11°55’ S, 14°51’E); restricted to the northern areas of Mombolo in the vicinity of Missão da Namba by Branch et al. 2021.  
Reproductionoviparous 
TypesHolotype: AMNH 47841, adult male 
DiagnosisDescription: Head and body dorsoventrally compressed; SVL 46.4–53.5 (mean 50.0) mm, HL 10.9– 12.6 (mean 11.6) mm, HW 7.8–11.2 (mean 10.6) mm, broadest at posterior level of eye and 1.1–1.4 (mean 1.3) times longer than wide. Eyes large (2.1–4.3 [mean 2.8] mm wide), pupil vertical with indented margins; circumorbital scales small and smooth, elongate at upper anterior margins, upper posterior scales with small upward pointing spines. Snout rounded, 4.5–5.0 (mean 4.3) mm long, slightly longer than distance between eye and ear openings (3.6–4.3 [mean 3.6] mm). Scales on top of snout smooth, rounded, scales to the side larger than central ones, with no intervening minute granules. Scales on snout slightly subequal in size to those on back of head or nape. Scales on eyelids larger than those on the crown. Circumorbital scales are separated by a row of smaller scales from the large scales on eyelid. Nostril pierced between rostral, two to three nasal scales and the 1st supralabial; the supranasal being much larger
than the subequal postnasals and being separated from each other by one to two smaller scales. Nostrils slightly elevated. Rostral roughly rectangular, but with its upper edges elongated due to extensions into the nostril. Supralabials 8–10 (mean 8.4). Infralabials 8–9 (mean 8.3). A total of 11–14 (mean 12.4) scales across the crown at level of front of eyes. A total of 14–16 (mean 15.4) scales from front of ear to back of eye. Mid-body scale rows 69–77 (mean 73.5). Original tail slightly dorsoventrally flattened and distinctly verticillate, with obvious lateral constrictions that are not that distinct to the tip of tail; each verticil comprising 5 imbricate rows of scales dorsally and 4 imbricate scale rows ventrally and with ventral scales approximately twice the size of those on the dorsal surface. Limbs well developed, hindlimbs slightly longer than forelimbs. All digits with a large pair of distal scansors, separated by a large, curved claw and followed after a large gap by a smaller pair of scansors; infero-median row of digital scales enlarged transversely, particularly towards the scansors, where the terminal scale adjoining the first pair of scansors may be medially constricted, swollen and scansor-like, enlarged subdigital lamellae on 4th toe 6–9 (mean 6.9). Precloacal pores 8 (Branch et al. 2021).

Colouration. In life (based on PEM R24187, Fig. 5A). Greyish above with six irregularly spaced darker grey crossbars from the occiput to the sacrum, central crossbars fused to form an X-shape; head with a dark grey band across the posterior edge of crown encompassing a small central pale spot; dark grey to black bar from nostril to the anterior margins of the ear opening; a vague, thin pale canthal stripe, extends on both sides from the nasal region to anterior margins of eye, continuing posteriorly of the eye to above the ear opening; upper and lower labials grey with diffuse yellow edges; lateral sides of the body with very faintly yellow-mustard scales scattered amongst more prominent darker grey scales; limbs greyish above with scattered darker grey markings with intervening faint yellow-mustard colouration; tail (regenerated) with irregular grey mottling; iris golden with a black narrow elliptic pupil with crenulated edge and black reticulation; ventre uniform greyish with scattered black specks; ventral limbs with scattered black specks, more prominent than on the ventrum. In preservative (based on PEM R24187): dorsum with six irregularly-spaced darker grey crossbars from the occiput to the sacrum with beige intervening blotches, central crossbars fused to from dark grey X-shape; ventre beige with numerous small scattered black specks on each scale, more prominent anteriorly, laterally and posteriorly. Variation: Greyish to brownish above with five to six irregularly-spaced darker grey-brown W-shaped crossbars from the occiput to the sacrum, sometimes fused in the middle to form X-shapes, limbs and tail with grey blotches; ventre uniform greyish with scattered black specks. Juveniles with more sharply-defined pattern. The original description of the holotype (Fig. 6) was based on a preserved and bleached specimen: ‘Above, greyish; back with five or six obsolescent, irregularly W-shaped brown crossbars; limbs and tail immaculate. Below, whitish, uniform’ (Loveridge 1944). Examination of images of the holotype confirm the presence of fine brown coloured specks ventrally that indicates that the ventre was not immaculate as originally described (Branch et al. 2021). 
CommentType species: Afroedura karroica bogerti LOVERIDGE 1944 is the type species of the genus Afroedura LOVERIDGE 1944.

Distribution: see map in Branch et al. 2017: 161, Branch et al. 2021: 57 (Fig. 1). 
EtymologyNamed after Charles Mitchill Bogert (1908-1992), former curator of the Department of Herpetology of the American Museum of Natural History. 
References
  • BARTS, M. & COLACICCO, F. 2019. Afrikanische Felsengeckos: Feldnotizen und Terrarienhaltung von Arten der Gattung Afroedura Loveridg e, 1944 – Teil III: Afroedura bogerti (Loveridg e, 1944). Sauria 41 (3): 31-36 - get paper here
  • Barts, M. & HAACKE, W. 2009. Afroedura bogerti (Loveridge, 1944) Reproduction. African Herp News (50): 39 - get paper here
  • Bauer, A.M. 2013. Geckos - The Animal Answer Guide. Johns Hopkins University Press, 159 pp.
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Branch WR, Haacke W, Vaz Pinto P, Conradie W, Baptista N, Verburgt L, Verisimmo L 2017. Loveridge’s Angolan geckos, Afroedura karroica bogerti and Pachydactylus scutatus angolensis (Sauria, Gekkonidae): new distribution records, comments on type localities and taxonomic status. Zoosystematics and Evolution 93(1): 157-166. - get paper here
  • Branch WR, Schmitz A, Lobón-Rovira J, Baptista NL, António T, Conradie W 2021. Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with descriptions of four new endemic species from Angola. Zoosystematics and Evolution 97(1): 55-82 - get paper here
  • BRANCH, WILLIAM R.; JENNIFER A. GUYTON, ANDREAS SCHMITZ, MICHAEL F. BAREJ, PIOTR NASKRECKI, HARITH FAROOQ, LUKE VERBURGT, MARK-OLIVER RÖDEL 2017. Description of a new flat gecko (Squamata: Gekkonidae: Afroedura) from Mount Gorongosa, Mozambique. Zootaxa 4324 (1): 142–160 - get paper here
  • Butler, B O; Ceríaco, L M P; Marques, M P; Bandeira, S; Júlio, T; Heinicke, M P & Bauer, A M; 2019. Herpetological survey of Huíla Province, Southwest Angola, including first records of Bicuar National Park. Herpetological Review 50 (2): 225-240 - get paper here
  • Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here
  • JACOBSEN, NIELS H. G.; ARIANNA L. KUHN, TODD R. JACKMAN & AARON M. BAUER 2014. A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa. Zootaxa 3846 (4): 451–501 - get paper here
  • Loveridge, A. 1944. New geckos of the genera Afroedura, new genus, and Pachydactylus from Angola. American Museum Novitates 1254: 1-4 - get paper here
  • Marques, Mariana P.; Luis M. P. Ceríaco , David C. Blackburn , and Aaron M. Bauer 2018. Diversity and Distribution of the Amphibians and Terrestrial Reptiles of Angola -- Atlas of Historical and Bibliographic Records (1840–2017). Proc. Cal. Acad. Sci. (Ser. 4) 65: 1-501 (Supplement II)
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
 
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