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Afroedura wulfhaackei BRANCH, SCHMITZ, LOBÓN-ROVIRA, BAPTISTA, ANTÓNIO & CONRADIE, 2021

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Angolan Flat Gecko 
SynonymAfroedura wulfhaackei BRANCH, SCHMITZ, LOBÓN-ROVIRA, BAPTISTA, ANTÓNIO & CONRADIE 2021: 66
Afroedura bogerti – BRANCH et al. 2017: 157 (part)
Afroedura bogerti – MARQUES et al. 2018: 177 (part)
Afroedura bogerti – BRANCH et al. 2019: 287 (part)
Afroedura bogerti (clade 4) – BRANCH et al. 2017: 147 
DistributionAngola (Benguela, N Huila, S Cuanza Sul, W Bie, Huambo)

Type locality: 3 km north of Maka-Mombolo (-12.17056, 14.88167, 1756 m a.s.l.), Benguela Province, Angola  
Reproduction 
TypesHolotype. PEM R24234, adult male, collected by William R. Branch and Pedro Vaz Pinto on 15 November 2016.
Paratypes. PEM R24232–3, adult females, collected 3 km north of Maka-Mombolo (-12.17056, 14.88167, 1756 m a.s.l), Benguela Province, Angola, by William R. Branch and Pedro Vaz Pinto on 15 November 2016; PEM R24236, adult male, collected above Maka-Mombolo, north-east of Balombo (-12.19833, 14.86833, 1857 m a.s.l.), Benguela Province, Angola, by William R. Branch and Pedro Vaz Pinto on 15 November 2016. 
DiagnosisDiagnosis. A member of the greater ‘transvaalica’ group in possessing two pairs of enlarged scansors per digit and a strongly verticillate and flattened tail (Jacobsen et al. 2014). It is part of the A. bogerti-group which differs from other members of the ‘transvaalica’ group by having less than 88 mid-body scale rows (vs. 97–102 in A. gorongosa, 113–120 in A. loveridgei, 102–119 in A. transvaalica); by the rostral bordering the nostril (nostril excluded from rostral in A. loveridgei); by the anterior nasals being mostly in contact ~ 68% (separated by 1–3 granules in A. gorongosa; always in broad contact in A. loveridgei; usually in broad contact in A. transvaalica ~ 3–18%); and in having 11–16 scales between the anterior borders of the eyes (19–22 in A. gorongosa; 15–19 in A. loveridgei; 15–20 in A. transvaalica) (comparative data fide Branch et al. 2017a).
Afroedura wulfhaackei sp. nov. differs from other members of the A. bogerti-group by a combination of the following characters (see Tables 3, 4): 76–88 (mean 79.3) mid-body scale rows (69–77 [mean 73.5] in A. bogerti, 64–78 [mean 72.8] in A. donveae sp. nov., 73–86 [mean 80.3] in A. vazpintorum sp. nov., 73–78 [mean 74.8] in A. praedicta sp. nov.); by the anterior nasals being mostly (~ 68% of the time) in contact (~ 33% of the time in contact in A. bogerti; always in contact in A. donveae sp. nov., A. vazpintorum sp. nov. and A. praedicta sp. nov.); each verticil comprising 4–5 (mean 4.0) ventral and 5–6 (mean 5.1) dorsal rows of scales (4 and 5 in A. bogerti and A. praedicta sp. nov.; 5–6 [mean 5.5] and 6–7 [mean 6.6] in A. donveae sp. nov.; 5–6 [mean 5.0] and 6–7 [mean 6.1] A. vazpintorum sp. nov.); ventral surfaces greyish with scattered small black spots (similar to A. bogerti and A. praedicta sp. nov., immaculate in A. donveae sp. nov. and A. vazpintorum sp. nov.). Afroedura wulfhaackei sp. nov. differs more specifically from its sister highland species A. bogerti in having a higher number of mid-body scale counts (76–88 [mean 79.3] versus 69–77 [mean 73.5]) and differs from A. praedicta sp. nov. in that the nasals are separated by smaller granules (versus always in contact) (Branch et al. 2021).

Colouration. In life (paratype PEM R24232, Fig. 5B): Greyish above with five irregularly-spaced darker crossbars from the occiput to the sacrum, each crossbar consisting posteriorly out of three to four black scales wide forming a W-shape; anterior to W-shape are 8–10 scales deep with a mix of dark grey and mustard colours; each dark crossbar separated by light grey to beige blotches; head with irregular dark grey-mustard blotches on the crown with intervening light grey colouration; dark mustard to dark grey bar from nostril to the anterior margins of the ear opening; a vague, thin pale grey canthal stripe, extends on both sides from the nasal region to anterior margins of eye; upper and lower labials grey with diffuse mustard edges; lateral sides of the body with a mix of dark grey and yellow-mustard colouration; limbs greyish above with scattered darker grey markings with intervening yellow-mustard colouration; tail (regenerated) with irregular grey-mustard mottling; iris golden with a black narrow elliptic pupil with crenulated edge and black reticulation with light grey intervening blotches; ventrum uniform greyish with scattered black specks; ventral limbs with scattered black specks, more prominent than on the ventrum. In preservative (holotype PEM R24234, Fig. 7): Dorsum with five irregularly-spaced dark grey W-shaped crossbars from the occiput to the sacrum with beige intervening blotches; ventrum is beige with numerous small scattered black specks on each scale, more prominent posteriorly. Variation: Greyish to brownish above with five to six irregularly-spaced darker grey-brown W-shaped crossbars from the occiput to the sacrum, limbs and tail with grey blotches; ventrum uniform greyish with scattered black specks. Juveniles have sharper patterns and colours (Branch et al. 2021). 
CommentDistribution: see map in Branch et al. 2021: 57 (Fig. 1). 
EtymologyThe new species is named in honour of Wulf Haacke, retired curator of the herpetology collection at the former Transvaal Museum (now Ditsong National Museum of Natural History). His herpetological expeditions to Angola in the early 1970s paved the way for this study and much of the material used in this study resulted from his expeditions. The name is constructed in the masculine singular genitive. 
References
  • Branch WR, Schmitz A, Lobón-Rovira J, Baptista NL, António T, Conradie W 2021. Rock island melody: A revision of the Afroedura bogerti Loveridge, 1944 group, with descriptions of four new endemic species from Angola. Zoosystematics and Evolution 97(1): 55-82 - get paper here
 
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