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Agama doriae BOULENGER, 1885

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards)
SubspeciesAgama doriae benueensis (MONARD 1951)
Agama doriae doriae BOULENGER 1885 
Common NamesE: Benoue Agama, Doria’s Agama, Nigeria Agama 
SynonymAgama doriae BOULENGER 1885: 127
Agama doriae — BOULENGER 1887: 495
Agama sennariensis WERNER 1914: 478 (fide MOODY & BÖHME 1984)
Agama kordofanensis WERNER 1919: 478 (fide  MOODY & BÖHME 1984)
Agama doriae cordofanensis WERNER 1919 (fide GEMEL et al. 2019)
Agama doriae — BARTS & WILMS 2003

Agama doriae benueensis (MONARD 1951)
Agama agama benueensis MONARD 1951: 131
Agama benueensis — GRANDISON 1968
Agama doriae benueensis — MOODY & BÖHME 1984
Agama doriae benueensis — WAGNER et al. 2009
Agama doriae benueensis — MEDIANNIKOV, TRAPE & TRAPE 2012
Agama doriae benueensis — TRAPE et al. 2012
Agama cf. benueensis — LEACHÉ et al. 2014 
DistributionGhana, Togo, Nigeria, Central African Republic to Eritrea and Ethiopia, N Cameroon, Sudan (Jumhūriyyat), Republic of South Sudan (RSS)

benueensis: Nigeria, Cameroon, Ghana; Type locality: Keren, Eritrea.

Type locality: Keren, pays des Bogos, Abyssinie septentrionale  
TypesTypes: BMNH (doriae)
Holotype: LCFM (also as MHNCF) [benueensis]
Syntypes: NMW [cordofanensis] 
CommentSynonymy: Has been previously synonymized with A. hartmanni by ANDERSON 1898.

Phylogenetics: Agama cf. benueensis is not closely related to A. doriae. It may be treated as a valid species. However, LEACHÉ et al. 2014 showed it as such in various trees without discussing its differences to it’s sister species, A. sankaranica and A. boensis.

Distribution: Agama doriae doesn’t occur in Mali. Not in Benin fide ULLENBRUCH et al. 2010. Map in Moody & Böhme 1984. A. d. benueensis is not in Nigeria fide MEDIANNIKOV et al. 2012. 
EtymologyNamed after Marchese Giacomo Doria (1840-1913), Italian zoologist who collected in Persia (Iran) with de Filippi (1862-1863) and in Borneo with Beccari (1865-1866). He founded the Museo Civico di Storia Naturale, Turin (1867-1913), and was its first Director. 
  • Anderson, J. 1898. Zoology of Egypt. Vol. l. Reptilia and Batrachia. B. Quaritch, London
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • Boulenger,G.A. 1885. Description d’une espèce nouvelle d’agame. Ann. Mus. Civ. Stor. Nat. Genova (2) 2: 127-128 - get paper here
  • Chirio, L. & Lebreton, M. 2007. Atlas des reptiles du Cameroun. MNHN, IRD, Paris 688 pp.
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Grandison, A. G. C. 1968. Nigerian lizards of the genus Agama (Sauria: Agamidae). Bull. Brit. Mus. Nat. Hist. (Zool.) 17: 65-90 - get paper here
  • Herrmann, Hans-Werner; Schmitz, Andreas; Herrmann, Patricia A. & Böhme, Wolfgang 2007. Amphibians and reptiles of the Tchabal Mbabo mountains, Adamaoua plateau, Cameroon. Bonner zoologische Beiträge 55 (1/2): 27-35 - get paper here
  • JACOBSEN, N.H.G. 2009. A contribution to the herpetofauna of the Passendro Area, Central African Republic. African Herp News (47): 2-20 - get paper here
  • Largen, M.J.; Spawls, S. 2006. Lizards of Ethiopia (Reptilia Sauria): an annotated checklist, bibliography, gazetteer and identification. Tropical Zoology 19 (1): 21-109 - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Leaché, A. et al. 2014. A hybrid phylogenetic–phylogenomic approach for species tree estimation in African Agama lizards with applications to biogeography, character evolution, and diversification. Molecular Phylogenetics and Evolution 79: 215-230, DOI: 10.1016/j.ympev.2014.06.013 - get paper here
  • LEACHÉ, Adam D.; Rebecca A. CHONG, Theodore J. PAPENFUSS, Philipp WAGNER, Wolfgang BÖHME, Andreas SCHMITZ, Mark-Oliver RÖDEL, Matthew LEBRETON, Ivan INEICH, Laurent CHIRIO, Aaron BAUER, Edem A. ENIANG, & Sherif BAHA EL DIN 2009. Phylogeny of the genus Agama based on mitochondrial DNA sequence data. Bonner zoologische Beiträge 56 (4): 273–278 - get paper here
  • Mediannikov, Oleg; Sébastien Trape, Jean-François Trape 2012. A Molecular Study of the Genus Agama (Squamata: Agamidae) in West Africa, with Description of Two New Species and a Review of the Taxonomy, Geographic Distribution, and Ecology of Currently Recognized Species. Russ. J. Herpetol. 19 (2): 115-142 - get paper here
  • Monard, A. 1951. Résultants de la mission zoologique suisse au Cameroun. Reptiles. Mém. IFAN, Dakkar, ser. sci. nat. 1: 123- 170.
  • Moody,S.M. & Böhme,W. 1984. Merkmalsvariation und taxonomische Stellung von Agama doriae Boulenger 1885 und Agama benueensis Monard 1951 (Reptilia: Agamidae) aus dem Sudangürtel Afrikas. Bonner Zoologische Beiträge 35: 107-128 - get paper here
  • Segniagbeto, Gabriel Hoinsoude; Jean-François Trape, Komlan M. Afiademanyo, Mark-Oliver Rödel, Annemarie Ohler, Alain Dubois, Patrick David, Danny Meirte, Isabelle Adolé Glitho, Fabio Petrozzi, and Luca Luiselli 2015. Checklist of the lizards of Togo (West Africa), with comments on systematics, distribution, ecology, and conservation. Zoosystema 37 (2): 381-402 - get paper here
  • Trape, J.F.; Trape, S. & Chirio, L. 2012. Lézards, crocodiles et tortues d'Afrique occidentale et du Sahara. IRD Orstom, 503 pp. - get paper here
  • Ullenbruch, K.; Grell, O.; Böhme, W. 2010. Reptiles from southern Benin, West Africa, with the description of a new Hemidactylus (Gekkonidae), and a country-wide checklist. Bonn Zool. Bull. 57 (1): 31-54 - get paper here
  • Wagner, P. 2014. A new cryptic species of the Agama lionotus complex from south of the Ngong Hills in Kenya. Salamandra 50 (4): 187-200 - get paper here
  • Wagner, P.; Barej, M.F. & Schmitz, A. 2009. Studies on African Agama VII. A new species of the Agama agama-group (Linnaeus, 1758) (Sauria: Agamidae) from Cameroon & Gabon, with comments on Agama mehelyi Tornier, 1902. Bonner zoologische Beiträge 56 (4): 285–297 - get paper here
  • Werner, F. 1914. Über Eidechsen, darunter zwei neue, aus dem angloägyptischen Sudan. Anz. Akad. Wissensch. Wien, ser. Mathemat.-Naturwissensch. 57 (18): 393-394
  • Werner,F. 1919. Wissenschaftliche Ergebnisse der mit Unterstützung der Kaiserlichen Akademie der Wissenschaften in Wien aus der Erbschaft Treitl von F. Werner unternommenen zoologischen Expedition nach dem Anglo-Aegyptischen Sudan (Kordofan) 1914. IV. Bearbeitung de Denkschr. Akad. Wiss Wien, Math.-Naturw. Klasse 96: 437-509 - get paper here
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