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Ahaetulla fusca (DUMÉRIL, BIBRON & DUMÉRIL, 1854)

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Higher TaxaColubridae, Ahaetuliinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Dark Whip Snake
G: Dunkler Baumschnüffler 
SynonymDryinus fuscus DUMÉRIL, BIBRON & DUMÉRIL 1854: 812
Dryophis mycterizans rhodogaster WALL 1908 (fide SMITH 1943)
Dryinus fuscus — DAVID & DUBOIS 2005
Dryophis mycterizans — BOULENGER 1896 (part) (nec Coluber mycterizans LINNAEUS 1758)
Dryophis mycterizans — WALL 1905 (part)
Dryophis mycterizans — BOURRET 1936
Dryophis mycterizans — DEUVE 1970
Passerita nasuta — COCHRAN 1930 (non Coluber nasutus LACÉPÈDE 1789)
Dryophis nasutus — SMITH 1943 (part) (nec Coluber nasutus LACÉPÈDE 1789)
Dryophis nasutus — Taylor 1965 (part)
Ahaetulla nasuta — STEJNEGER 1933 (part) (nec Coluber nasutus LACÉPÈDE 1789)
Ahaetulla nasuta — DOWLING & JENNER 1988
Ahaetulla nasuta — COX 1991
Ahaetulla nasuta — VOGEL 1992
Ahaetulla nasuta — MANTHEY & GROSSMANN 1997 (part)
Ahaetulla nasuta — COX et al. 1998
Ahaetulla nasuta — COX et al. 2012
Ahaetulla nasuta — ZUG et al. 1998
Ahaetulla nasuta — CHAN-ARD et al. 1999
Ahaetulla nasuta — HALLERMANN et al. 2002 (part)
Ahaetulla nasuta — NGUYEN et al. 2009
Ahaetulla nasuta — GRISMER et al. 2007
Ahaetulla nasuta — GRISMER et al. 2008
Ahaetulla nasuta — WALLACH et al. 2014 (part)
Ahaetulla nasuta — ZUG et al. 1998
Dryophis mycterizans var. rhodogaster WALL 1908: 919
Dryophis mycterizans var. rhodogaster — WALL 1910
Dryophis mycterizans rhodogaster — SMITH 1943
Ahaetulla fusca — DAVID et al. 2022 
DistributionThailand, Cambodia, S. Vietnam, Myanmar

Type locality: “Bangkok”, Thailand (neotype locality)

rhodogaster: Type locality. “Shwebo, Upper Burma,” now Shwebo, Sagaing Region (22.566 °N 95.683 °E), Myanmar.  
Reproductionovoviviparous (5-15 live offspring) 
TypesNeotype. BMNH 1956.1.12.99, an adult male, collected in 1946 by a Thai assistant of Dr. M. T. Gillies and presented by J. D. Romer, by designation of David and Dubois (2005: 173).
Holotype. BNHS 1982, an adult female obtained from Major H. Delmé-Radcliffe, 2 July 1908, fide David et al. 2022 [rhodogaster] 
DiagnosisDiagnosis: “A large species of the genus Ahaetulla inhabiting Myanmar (? See below), Thailand, Laos, Cambodia and Vietnam, characterized by the following combination of characters: (1) snout acuminate, slightly convex above, not depressed, terminating in a rostral elongate and shaped as a long, pointed dermal appendage; (2) a long rostral appendage, slightly curved downward, and shorter than horizontal eye diameter (vs. short rostral appendage in A. isabellina, A. malabarica. A. farnsworthi, A. borealis, and vs. no rostral appendage in A. mycterizans, A. prasina, A. fronticincta, A. dispar, A. travancorica, and A. perroteti); (3) rostral appendage composed of a single grooved, elongate scale (vs. rostral appendage composed of multiple small scales in A. pulverulenta, A. sahyadrensis, and A. anomala); (4) loreal scale usually absent, internasals and prefrontals in contact with supralabials but, sometimes, 1 large loreal resulting from the longitudinal division of the 3rd supralabial (vs. always present in A. dispar, A. travancorica); (5) usually 1 anterior temporal; (6) body usually an overall bright or deep green coloration, sometimes gray, buff, yellow, tan or pale brown (vs. usually gray-brown in A. pulverulenta, A. sahyadrensis and often brown in adult females of A. anomala); (7) no crown markings (vs. a distinct rhomboid crown marking present on top of head in A. sahyadrensis and A. anomala); (8) cloacal plate divided; (9) venter uniform with a pale, white or yellow longitudinal stripe on each outer edge of ventral plates; (10) at least 174 ventral plates.” (David et al. 2022)


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CommentSynonymy: after David et al. 2022 but not exhaustive. See David et al. 2022 for sources. Previsously considered as a synonym of A. nasutus but revalidated by David et al. 2022.

Distribution: See David et al. 2022: 25 (Fig. 1) for a map.

Habitat: fully arboreal (by implication, Harrington et al. 2018).

Types: Dryinus fuscus was described from two syntypes representing two distinct species, one being MNHN 7565, now lost, alleged to be from “Java.” David and Dubois (2005) showed that specimen MNHN 7565 is, in fact, the holotype of Ahaetulla pulverulenta. See David et al. 2022 for further discussion. 
EtymologyFrom Latin, nasuta = ‘of the nose’ an adjective alluding to its elongate snout. 
References
  • David, Patrick; Bartoz Nadolski, Sumaithangi Rajagopalan Ganesh, Omkar Dilip Adhikari, Achyuthan N. Srikanthan 2022. A Nomenclatural Review of Ahaetulla nasuta (Lacépède, 1789): resurrection of Ahaetulla fusca (Duméril, Bibron et Duméril, 1854) for the populations of the Indochinese Region (Reptilia, Squamata, Colubridae). Russian Journal of Herpetology 29 (1): 19–34. - get paper here
  • Duméril, A. M. C., Bibron, G. & DUMÉRIL, A. H. A., 1854. Erpétologie générale ou histoire naturelle complète des reptiles. Tome septième. Deuxième partie, comprenant l'histoire des serpents venimeux. Paris, Librairie Encyclopédique de Roret: i-xii + 781-1536 - get paper here
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
 
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