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Amphiesma stolatum (LINNAEUS, 1758)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Buff Striped Keelback
G: Gelbhalswassernatter, Gestreifte Wassernatter, Asiatische Streifennatter, Gelbband-Wassernatter
Chinese: 草腹链蛇 
SynonymColuber stolatus LINNAEUS 1758: 219
Elaps bilineatus SCHNEIDER 1801: 299
Natrix stolatus — MERREM 1820: 123
Tropidonotus stolatus — BOIE 1827: 535
Amphiesma stolatum DUMÉRIL, BIBRON & DUMÉRIL 1854: 727
Tropidonotus stolbatus [sic] — KELAART 1854: 139 (in error ?)
Tropidonotus stolatus — GÜNTHER 1864: 266
Tropidonotus ruficeps PETERS 1869: 444
Tropidonotus stolatus — STOLICZKA 1870: 191
Tropidonotus stolatus — BOULENGER 1890: 348
Tropidonotus stolatus — BOULENGER 1893: 253
Amphiesma stolatum — COPE 1895: 425
Natrix stolata — STEJNEGER 1907: 280
Tropidonotus stolatus — WALL 1908: 320
Tropidonotus stolatus — WALL 1919: 562
Rhabdophis stolatus — WALL 1921
Amphiesma stolata — WALL 1921: 105
Natrix stolata chinensis MELL 1930
Rhabdophis stolatus chinensis — BOURRET 1935: 4
Natrix stolata chinensis — RENDAHL 1937
Natrix stolata — SMITH 1943: 303
Rhabdophis stolatus var. chinensis — DEUVE 1961
Amphiesma stolata — DAS 1996: 53
Amphiesma stolata — COX et al. 1998: 45
Amphiesma stolatum — DAVID, VOGEL & PAUWELS 1998
Amphiesma stolata chinesis — CHAN-ARD et al. 1999: 154
Amphiesma stolata — SHARMA 2004
Amphiesma stolata — JANZEN et al. 2007
Amphiesma stolatus — GUO et al. 2014
Amphiesma stolatum — WALLACH et al. 2014: 34
Amphiesma solatum — NEPALI & SINGH 2022 (in error) 
DistributionTaiwan (incl. Lanyu),
China (Hainan, Hong Kong, Fujian, Jiangxi),
Sri Lanka, Andaman Island ?,
India (Kerala, Tamil Nadu, Andhra Pradesh, Karnataka, Gujarat, Madhya Pradesh, Chhattisgarh, Orissa = Odisha, Uttar Pradesh, Assam, Bihar, Maharashtra (Koyna, Marunji, Lohagad, Lonavla, Phansad), Arunachal Pradesh (Miao - Changlang district, Chimpu, Van Vihar, Chessa - Papum Pare district), Punjab [Dino Aulakh, pers. comm.], Himachal Pradesh, Jammu and Kashmir, Mizoram, Telangana, Nagaland), Bhutan,
Myanmar (= Burma), Thailand, Vietnam (Hoa Binh), Laos, Cambodia, Nepal, Pakistan

Type locality: “America” (in error); “Indien” fide KRAMER 1977.  
Reproductionoviparous 
TypesHolotype: NRM (Stockholm) 
DiagnosisDiagnosis (genus). Solid-toothed snakes with keeled scales on the body in 19 dorsal rows mid-body, anal usually divided, and all subcaudals divided. Everted hemipenis is Y shape with simple sulcus (from Guo et al. 2014).
Description: “Eye moderate. Rostral twice as broad as deep, just visible from above; internasals much narrowed anteriorly, as long as broad or a little longer, nearly as long as the prefrontals frontal once and two thirds to twice as long as broad, longer than its distance from the end of the snout, as long as the parietals; loreal as long as deep or deeper one prae- and three postoculars; temporals 1+1 or 1+2; eight upper labials normally, third, fourth, and fifth entering the eye; five or six lower labials in contact with the anterior chin-shields, which are shorter than the posterior. Scales in 19 rows, strongly keeled, outer row smooth. Ventral 120-161; anal divided : subcaudals 50-89. Greenish or brownish olive, with more or less distinct black spots or reticulated cross bars intersected by two yellow longitudinal bands, which are best marked posteriorly; præ- and postoculars yellowish lower surfaces white, usually with a black spot on each side of each ventral.” (Boulenger 1893: 253) 
CommentSynonymy partly after STEJNEGER 1907. Kaiser et al. 2013 considered the generic names Asianatrix Hoser 2012, Elliottnatrix Hoser 2012, Greernatrix Hoser 2012, Sundanatrix Hoser 2012, Wellingtonnatrix Hoser 2012, Wellsnatrix Hoser 2012 invalid and rejected their use instead of Amphiesma.

Type species: Coluber stolatus LINNAEUS 1758 is the type species of the genus Amphiesma DUMÉRIL, BIBRON & DUMÉRIL 1854 (see DAVID et al. 1998).

Guo et al. 2014 restricted the genus Amphiesma to A. stolatum, moving all the other species to Hebius and Herpetoreas.

Distribution: Has been reported from the Andaman Islands by SMITH 1943 and others although no voucher specimens exist. The existence on these islands is therefore questionable (Das 1999). STUEBING (1991) reported this species from Borneo but this appears to be based on a misidentification according to MALKMUS et al. (2002). For a map see Sindaco et al. 2013.

Behavior: diurnal. 
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